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Hemichordates ("half chordates") have some features similar to those of chordates: branchial openings that open into the pharynx and look rather like gill slits; stomochords, similar in composition to notochords, but running in a circle round the "collar", which is ahead of the mouth; and a dorsal nerve cord—but also a smaller ventral nerve cord.
The Cambrian chordates are characterised by the presence of segmented muscle blocks called myomeres and notochord, the two defining features of chordates. Before the full understanding of Cambrian fossils, chordates as members the most advanced phylum were believed to appear on Earth much later than the Cambrian.
Stylophorans are classed as echinoderms based on their possession of at least two shared and unique features (apomorphies) of the phylum; stereom plates and a water vascular system. Because they show no sign of radial symmetry, the position of stylophorans within Echinodermata remains unresolved as of 2019. [2]
Lancelets diverged from other chordates during or prior to the Cambrian period. A number of fossil chordates have been suggested to be closely related to lancelets, including Pikaia and Cathaymyrus from the Cambrian and Palaeobranchiostoma from the Permian, but their close relationship to lancelets has been doubted by other authors.
A tunicate is an exclusively marine invertebrate animal, a member of the subphylum Tunicata (/ ˌ tj uː n ɪ ˈ k eɪ t ə / TEW-nih-KAY-tə).This grouping is part of the Chordata, a phylum which includes all animals with dorsal nerve cords and notochords (including vertebrates).
Pikaia gracilens is an extinct, primitive chordate marine animal known from the Middle Cambrian Burgess Shale of British Columbia.Described in 1911 by Charles Doolittle Walcott as an annelid, and in 1979 by Harry B. Whittington and Simon Conway Morris as a chordate, it became "the most famous early chordate fossil", [1] or "famously known as the earliest described Cambrian chordate". [2]
The dorsal nerve cord is an anatomical feature found in chordate animals, mainly in the subphyla Vertebrata and Cephalochordata, as well as in some hemichordates.It is one of the five embryonic features unique to all chordates, the other four being a notochord, a post-anal tail, an endostyle, and pharyngeal slits.
Netrin is important in patterning of the neural system in chordates, as well as is the molecule Shh, but S. kowalevskii was only found to have one hh gene and it appears to be expressed in a region that is uncommon to where it is usually expressed in developing chordates along the ventral midline.