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Horizontal symbiont transfer (horizontal transmission) is a process where a host acquires a facultative symbiont from the environment or another host. [9] The Rhizobia-Legume symbiosis (bacteria-plant endosymbiosis) is a prime example of this modality. [21]
Secondary endosymbiosis has occurred several times and has given rise to extremely diverse groups of algae and other eukaryotes. Some organisms can take opportunistic advantage of a similar process, where they engulf an alga and use the products of its photosynthesis, but once the prey item dies (or is lost) the host returns to a free living state.
The theory of endosymbiosis, as known as symbiogenesis, provides an explanation for the evolution of eukaryotic organisms. According to the theory of endosymbiosis for the origin of eukaryotic cells, scientists believe that eukaryotes originated from the relationship between two or more prokaryotic cells approximately 2.7 billion years ago.
Diagram of the six possible types of symbiotic relationship, from mutual benefit to mutual harm. The definition of symbiosis was a matter of debate for 130 years. [7] In 1877, Albert Bernhard Frank used the term symbiosis to describe the mutualistic relationship in lichens.
Evidence for primary endosymbiosis includes the presence of a double membrane around the chloroplasts; one membrane belonged to the bacterium, and the other to the eukaryote that captured it. Over time, many genes from the chloroplast have been transferred to the nucleus of the host cell through endosymbiotic gene transfer (EGT).
Reductive evolution is the process by which microorganisms remove genes from their genome.It can occur when bacteria found in a free-living state enter a restrictive state (either as endosymbionts or parasites) or are completely absorbed by another organism becoming intracellular (symbiogenesis).
Secondary endosymbiosis results in the engulfment of an organism that has already performed primary endosymbiosis. Thus, four plasma membranes are formed. The first originating from the cyanobacteria, the second from the eukaryote that engulfed the cyanobacteria, and the third from the eukaryote who engulfed the primary endosymbiotic eukaryote. [11]
Endogenosymbiosis is an evolutionary process, proposed by the evolutionary and environmental biologist Roberto Cazzolla Gatti, in which "gene carriers" (viruses, retroviruses and bacteriophages) and symbiotic prokaryotic cells (bacteria or archaea) could share parts or all of their genomes in an endogenous symbiotic relationship with their hosts.