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The term Monera became well established in the 20s and 30s when to rightfully increase the importance of the difference between species with a nucleus and without. In 1925, Édouard Chatton divided all living organisms into two sections, Prokaryotes and Eukaryotes: the Kingdom Monera being the sole member of the Prokaryotes section. [23]
Prokaryotes evolved before eukaryotes, and lack nuclei, mitochondria, and most of the other distinct organelles that characterize the eukaryotic cell. Some unicellular prokaryotes, such as cyanobacteria , form colonies held together by biofilms , and large colonies can create multilayered microbial mats .
[3] [4] In the three-domain model, the first two are prokaryotes, single-celled microorganisms without a membrane-bound nucleus. All organisms that have a cell nucleus and other membrane-bound organelles are included in Eukarya and called eukaryotes. Non-cellular life, most notably the viruses, is not included in this system.
The three-domain system adds a level of classification (the domains) "above" the kingdoms present in the previously used five- or six-kingdom systems.This classification system recognizes the fundamental divide between the two prokaryotic groups, insofar as Archaea appear to be more closely related to eukaryotes than they are to other prokaryotes – bacteria-like organisms with no cell nucleus.
Traditionally, Archaea only included its prokaryotic members, but this since has been found to be paraphyletic, as eukaryotes are now known to have evolved from archaea. Even though the domain Archaea includes eukaryotes, the term "archaea" ( sg. : archaeon / ɑːr ˈ k iː ɒ n / ar- KEE -on , from the Greek "ἀρχαῖον", which means ...
They held that only monophyletic groups should be accepted as formal ranks in a classification and that – while this approach had been impractical previously (necessitating "literally dozens of eukaryotic 'kingdoms '") – it had now become possible to divide the eukaryotes into "just a few major groups that are probably all monophyletic". [42]
For the Prokaryotes (Bacteria and Archaea) the rank kingdom has not been used till 2024 [115] (although some authors referred to phyla as kingdoms [73]). The category of kingdom was included into the Bacteriological Code in November 2023, [ 116 ] the first four proposals ( Bacillati , Fusobacteriati , Pseudomonadati , Thermotogati ) were ...
Phylogenetic and symbiogenetic tree of living organisms, showing the origins of eukaryotes and prokaryotes. The two-empire system (two-superkingdom system) was the top-level biological classification system in general use from the early 20th century until the establishment of the three-domain system (which itself is currently being challenged by the two-domain system).