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This membrane contains an enzyme called NAD(P)H dehydrogenase which transfers electrons in a linear chain to oxygen molecules. [1] This electron transport chain (ETC) within the chloroplast also interacts with those in the mitochondria where respiration takes place. [2] Photosynthesis is also a process that Chlororespiration interacts with. [2]
Cyanobacteria is the only prokaryotic group that performs oxygenic photosynthesis. Anoxygenic photosynthetic bacteria use PSI- and PSII-like photosystems, which are pigment protein complexes for capturing light. [5] Both of these photosystems use bacteriochlorophyll. There are multiple hypotheses for how oxygenic photosynthesis evolved.
The reaction center is in the thylakoid membrane. It transfers absorbed light energy to a dimer of chlorophyll pigment molecules near the periplasmic (or thylakoid lumen) side of the membrane. This dimer is called a special pair because of its fundamental role in photosynthesis. This special pair is slightly different in PSI and PSII reaction ...
The reaction begins with the excitation of a pair of chlorophyll molecules similar to those in the bacterial reaction center. Due to the presence of chlorophyll a, as opposed to bacteriochlorophyll, Photosystem II absorbs light at a shorter wavelength. The pair of chlorophyll molecules at the reaction center are often referred to as P680. [1]
Although the functions of photorespiration remain controversial, [20] it is widely accepted that this pathway influences a wide range of processes from bioenergetics, photosystem II function, and carbon metabolism to nitrogen assimilation and respiration.
The organic or inorganic substances (e.g., oxygen) used as electron acceptors needed in the catabolic processes of aerobic or anaerobic respiration and fermentation are not taken into account here. For example, plants are lithotrophs because they use water as their electron donor for the electron transport chain across the thylakoid membrane.
The antenna complex contains hundreds of chlorophyll molecules which funnel the excitation energy to the center of the photosystem. At the reaction center, the energy will be trapped and transferred to produce a high energy molecule. [2] The main function of PSII is to efficiently split water into oxygen molecules and protons.
Photoheterotrophs generate ATP using light, in one of two ways: [6] [7] they use a bacteriochlorophyll-based reaction center, or they use a bacteriorhodopsin.The chlorophyll-based mechanism is similar to that used in photosynthesis, where light excites the molecules in a reaction center and causes a flow of electrons through an electron transport chain (ETS).