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Archaea and bacteria have generally similar cell structure, but cell composition and organization set the archaea apart. Like bacteria, archaea lack interior membranes and organelles. [68] Like bacteria, the cell membranes of archaea are usually bounded by a cell wall and they swim using one or more flagella. [119]
The two-domain system is a biological classification by which all organisms in the tree of life are classified into two domains, Bacteria and Archaea. [1] [2] [3] It emerged from development of knowledge of archaea diversity and challenges the widely accepted three-domain system that classifies life into Bacteria, Archaea, and Eukarya. [4]
Phylogenetic tree showing the relationship between the archaea and other forms of life. Eukaryotes are colored red, archaea green and bacteria blue. Adapted from Ciccarelli et al. [44] Woese argued that the bacteria, archaea, and eukaryotes represent separate lines of descent that diverged early on from an ancestral colony of organisms.
The three-domain system adds a level of classification (the domains) "above" the kingdoms present in the previously used five- or six-kingdom systems.This classification system recognizes the fundamental divide between the two prokaryotic groups, insofar as Archaea appear to be more closely related to eukaryotes than they are to other prokaryotes – bacteria-like organisms with no cell nucleus.
In 1977, Carl Woese proposed dividing prokaryotes into the Bacteria and Archaea (originally Eubacteria and Archaebacteria) because of the major differences in the structure and genetics between the two groups of organisms. Archaea were originally thought to be extremophiles, living only in inhospitable conditions such as extremes of temperature ...
A speculatively rooted tree for RNA genes, showing major branches Bacteria, Archaea, and Eukaryota The three-domain tree and the eocyte hypothesis (two-domain tree), 2008. [7] Phylogenetic tree showing the relationship between the eukaryotes and other forms of life, 2006. [8] Eukaryotes are colored red, archaea green, and bacteria blue.
Halophiles, grow optimally in high salt concentrations between 0.2 M and 5.2 M NaCl, including the genera Haloarcula, Haloferax, Halococcus. [23] [26] Methanogens are a significant subset of archaea and include many extremophiles, but are also ubiquitous in wetland environments as well as the ruminant and hindgut of animals. [27]
Archaea share this defining feature with the bacteria with which they were once grouped. In 1990 the microbiologist Woese proposed the three-domain system that divided living things into bacteria, archaea and eukaryotes, [42] and thereby split the prokaryote domain. Archaea differ from bacteria in both their genetics and biochemistry.