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In the stems of some Asterales dicots, there may be phloem located inwardly from the xylem as well. Between the xylem and phloem is a meristem called the vascular cambium. This tissue divides off cells that will become additional xylem and phloem. This growth increases the girth of the plant, rather than its length. As long as the vascular ...
The cells of the vascular cambium (F) divide to form phloem on the outside, located beneath the bundle cap (E), and xylem (D) on the inside. Most of the vascular cambium is here in vascular bundles (ovals of phloem and xylem together) but it is starting to join these up as at point F between the bundles.
The movement in phloem is multidirectional, whereas, in xylem cells, it is unidirectional (upward). [citation needed] [15] After the growth period, when the meristems are dormant, the leaves are sources, and storage organs are sinks. Developing seed-bearing organs (such as fruit) are always sinks. Because of this multi-directional flow, coupled ...
The presence of vessels in xylem has been considered to be one of the key innovations that led to the success of the flowering plants. It was once thought that vessel elements were an evolutionary innovation of flowering plants, but their absence from some basal angiosperms and their presence in some members of the Gnetales suggest that this hypothesis must be re-examined; vessel elements in ...
There is also a tissue between xylem and phloem, which is the cambium. The xylem typically lies towards the axis ( adaxial ) with phloem positioned away from the axis ( abaxial ). In a stem or root this means that the xylem is closer to the centre of the stem or root while the phloem is closer to the exterior.
The fibers of the xylem are always lignified, while those of the phloem are cellulosic. Reliable evidence for the fibre cells' evolutionary origin from tracheids exists. [ 10 ] During evolution the strength of the tracheid cell walls was enhanced, the ability to conduct water was lost and the size of the pits was reduced.
In a vascular plant, the stele is the central part of the root or stem [1] containing the tissues derived from the procambium. These include vascular tissue, in some cases ground tissue and a pericycle, which, if present, defines the outermost boundary of the stele.
Tracheids were the main conductive cells found in early vascular plants. In the first 140–150 million years of vascular plant evolution, tracheids were the only type of conductive cells found in fossils of plant xylem tissues. [5] Ancestral tracheids did not contribute significantly to structural support, as can be seen in extant ferns. [6]