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However, the punctuational equilibrium model may still be inferred from both the observation of stasis and examples of rapid and episodic speciation events documented in the fossil record. [70] Dawkins also emphasizes that punctuated equilibrium has been "oversold by some journalists", [71] but partly due to Eldredge and Gould's "later writings ...
There are four geographic modes of speciation in nature, based on the extent to which speciating populations are isolated from one another: allopatric, peripatric, parapatric, and sympatric. Whether genetic drift is a minor or major contributor to speciation is the subject of much ongoing discussion.
In the 1980s, the American palaeontologists Stephen Jay Gould and Niles Eldredge argued for an extended synthesis based on their idea of punctuated equilibrium, the role of species selection shaping large scale evolutionary patterns and natural selection working on multiple levels extending from genes to species. [30] [31] [32] [33]
Punctuated gradualism is considered to be a variation of these models, lying somewhere in between the phyletic gradualism model and the punctuated equilibrium model. It states that speciation is not needed for a lineage to rapidly evolve from one equilibrium to another but may show rapid transitions between long-stable states.
The Bak–Sneppen model is a simple model of co-evolution between interacting species. It was developed to show how self-organized criticality may explain key features of the fossil record, such as the distribution of sizes of extinction events and the phenomenon of punctuated equilibrium. It is named after Per Bak and Kim Sneppen.
Gould's most significant contribution to evolutionary biology was the theory of punctuated equilibrium [2] developed with Niles Eldredge in 1972. [3] The theory proposes that most evolution is characterized by long periods of evolutionary stability, infrequently punctuated by swift periods of branching speciation.
A period of stasis then occurs until the species either becomes extinct or splits again. Gould argued that punctuated equilibrium challenges the gradual change expected by extrapolationists. In the case of Hominid evolution, there is the evolutionary trend of marked increase in brain size.
This can lead to the opposite occurring with disruptive selection eventually selecting against the average; when everyone starts taking advantage of that resource it will become depleted and the extremes will be favored. Furthermore, gradualism is a more realistic view when looking at speciation as compared to punctuated equilibrium.