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The light excites an electron in the pigment P680 at the core of photosystem II, which is transferred to the primary electron acceptor, pheophytin, leaving behind P680 +. The energy of P680 + is used in two steps to split a water molecule into 2H + + 1/2 O 2 + 2e - ( photolysis or light-splitting ).
This chain of electron acceptors is known as an electron transport chain. When this chain reaches PSI, an electron is again excited, creating a high redox-potential. The electron transport chain of photosynthesis is often put in a diagram called the Z-scheme, because the redox diagram from P680 to P700 resembles the letter Z. [3]
At the reaction center, the electrons on the special chlorophyll molecule will be excited and ultimately transferred away by electron carriers. (If the electrons were not transferred away after excitation to a high energy state, they would lose energy by fluorescence back to the ground state, which would not allow plants to drive photosynthesis.)
P680 + is the strongest biological oxidizing agent known, with an estimated redox potential of ~1.3 V. [3] This makes it possible to oxidize water during oxygenic photosynthesis. P680 + recovers its lost electron by oxidizing water via the oxygen-evolving complex , which regenerates P680.
Photosystem II (or water-plastoquinone oxidoreductase) is the first protein complex in the light-dependent reactions of oxygenic photosynthesis. It is located in the thylakoid membrane of plants , algae , and cyanobacteria .
This membrane contains an enzyme called NAD(P)H dehydrogenase which transfers electrons in a linear chain to oxygen molecules. [1] This electron transport chain (ETC) within the chloroplast also interacts with those in the mitochondria where respiration takes place. [2] Photosynthesis is also a process that Chlororespiration interacts with. [2]
It is possible to introduce an artificial electron acceptor into the light reaction, such as a dye that changes color when it is reduced. These are known as Hill reagents. These dyes permitted the finding of electron transport chains during photosynthesis. Dichlorophenolindophenol (DCPIP), an example of these dyes, is widely used by experimenters.
Photosynthesis and cellular respiration are distinct processes, as they take place through different sequences of chemical reactions and in different cellular compartments (cellular respiration in mitochondria). [15] [16] The general equation for photosynthesis as first proposed by Cornelis van Niel is: [17]
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