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A chordate (/ ˈ k ɔːr d eɪ t / KOR-dayt) is a deuterostomal bilaterian animal belonging to the phylum Chordata (/ k ɔːr ˈ d eɪ t ə / kor-DAY-tə).All chordates possess, at some point during their larval or adult stages, five distinctive physical characteristics (synapomorphies) that distinguish them from other taxa.
Idealised vertebrate body plan, showing key characteristics. Vertebrates (and other chordates) belong to the Bilateria, a group of animals with mirror symmetrical bodies. [6] They move, typically by swimming, using muscles along the back, supported by a strong but flexible skeletal structure, the spine or vertebral column. [7]
On the other hand, fossils of early chordates are very rare, as non-vertebrate chordates have no bone tissue or teeth, and fossils of no Post-Cambrian non-vertebrate chordates are known aside from the Permian-aged Paleobranchiostoma, trace fossils of the Ordovician colonial tunicate Catellocaula, and various Jurassic-aged and Tertiary-aged ...
Despite their simple appearance and very different adult form, their close relationship to the vertebrates is certain. Both groups are chordates, as evidenced by the fact that during their mobile larval stage, tunicates possess a notochord, a hollow dorsal nerve cord, pharyngeal slits, post-anal tail, and an endostyle. They resemble a tadpole.
The notochord is the defining feature (synapomorphy) of chordates, and was present throughout life in many of the earliest chordates. Although the stomochord of hemichordates was once thought to be homologous or from a common lineal origin, it is now viewed as analogous, convergent , or from a different lineal origin. [ 22 ]
According to this theory, there is a deep agreement between the vertebrates and cephalochordates, and even all chordates. [75] [76] With the advent of molecular genetics lancelets are once again regarded as a model of vertebrate ancestors, and are used again as a model organism. [77] [30]
This, combined with an apparent lack of vertebral elements within the Myxini, suggested that the Myxini were descended from a more ancient lineage than the vertebrates, and that the skull developed before the vertebral column. The clade was thus composed of the Myxini and the vertebrates, and any extinct chordates with skulls.
However, it is now accepted [who?] that it is the vertebrate pharyngeal pouches and not the neck slits that are homologous to the pharyngeal slits of invertebrate chordates. [citation needed] Pharyngeal arches, pouches, and clefts are, at some stage of life, found in all chordates. One theory of their origin is the fusion of nephridia which ...