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The basic function of the xylem is to transport water upward from the roots to parts of the plants such as stems and leaves, but it also transports nutrients. [1] [2] The word xylem is derived from the Ancient Greek word, ξύλον (xylon), meaning "wood"; the best-known xylem tissue is wood, though it is found throughout a plant. [3]
It is used mainly for the transport of water, along with some small nutrients. The meristem of the stem creates cells which make up the cambium and pro-cambium. These cells then produce a highly branched poly-phenolic protein, called lignin, in a very high concentration. The cells then perform apoptosis, and the actual xylem tube begins to form ...
The cells in vascular tissue are typically long and slender. Since the xylem and phloem function in the conduction of water, minerals, and nutrients throughout the plant, it is not surprising that their form should be similar to pipes. The individual cells of phloem are connected end-to-end, just as the sections of a pipe might be.
Stained Xylem Vessel in Red/Orange (x400 magnification) In secondary xylem – the xylem that is produced as a stem thickens rather than when it first appears – a vessel element originates from the vascular cambium. A long cell, oriented along the axis of the stem, called a "fusiform initial", divides along its length forming new vessel elements.
With much of the sucrose having been removed, the water exits the phloem by osmosis or is drawn by transpiration into nearby xylem vessels, lowering the turgor pressure within the phloem. [4] The sucrose concentration in sieve tubes is typically 10–30% in the leaves but only 0.5% in the photosynthesis cells.
The cells of the vascular cambium (F) divide to form phloem on the outside, located beneath the bundle cap (E), and xylem (D) on the inside. Most of the vascular cambium is here in vascular bundles (ovals of phloem and xylem together) but it is starting to join these up as at point F between the bundles.
It is a type of conductive cell called a tracheary element. Angiosperms also use another type of conductive cell, called vessel elements, to transport water through the xylem. The main functions of tracheid cells are to transport water and inorganic salts, and to provide structural support for trees.
Its most commonly noted function is the support through strengthening of wood (mainly composed of xylem cells and lignified sclerenchyma fibres) in vascular plants. [17] [18] [19] Finally, lignin also confers disease resistance by accumulating at the site of pathogen infiltration, making the plant cell less accessible to cell wall degradation. [20]