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Kinocilia are present in the crista ampullaris of the semicircular ducts and the sensory maculae of the utricle and saccule. [1] One kinocilium is the longest cilium located on the hair cell next to 40–70 stereocilia. During movement of the body, the hair cell is depolarized when the stereocilia move toward the kinocilium.
Stereocilia (or stereovilli or villi) are non-motile apical cell modifications. They are distinct from cilia and microvilli , but are closely related to microvilli. They form single "finger-like" projections that may be branched, with normal cell membrane characteristics.
Each hair cell of a macula has 40 to 70 stereocilia and one true cilium called a kinocilium. The stereocilia are oriented by the striola, a curved ridge that runs through the middle of the macula; in the saccule they are oriented away from the striola [2] The tips of the stereocilia and kinocilium are embedded in a gelatinous otolithic membrane ...
The kinocilium is the only sensory aspect of the hair cell and is what causes hair cell polarization. The tips of these stereocilia and kinocilium are embedded in a gelatinous layer, which together with the statoconia form the otolithic membrane. [2] This membrane is weighted with calcium carbonate-protein granules called otoliths.
Resembling hair-like projections, the stereocilia are arranged in bundles of 30–300. [3] Within the bundles the stereocilia are often lined up in several rows of increasing height, similar to a staircase. At the core of these hair-like stereocilia are rigid cross-linked actin filaments, which can renew every
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As a result, the cupula is deflected opposite the direction of head movement. As the endolymph pushes the cupula, the stereocilia is bent as well, stimulating the hair cells within the crista ampullaris. After a short time of continual rotation however, the endolymph's acceleration normalizes with the rate of rotation of the semicircular ducts.
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