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Stroma, in botany, refers to the colorless fluid surrounding the grana within the chloroplast. [ 1 ] Within the stroma are grana (stacks of thylakoid ), the sub-organelles where photosynthesis is started [ 2 ] before the chemical changes are completed in the stroma.
Within the envelope membranes, in the region called the stroma, there is a system of interconnecting flattened membrane compartments, called the thylakoids.The thylakoid membrane is quite similar in lipid composition to the inner envelope membrane, containing 78% galactolipids, 15.5% phospholipids and 6.5% sulfolipids in spinach chloroplasts. [3]
The inner chloroplast membrane borders the stroma and regulates passage of materials in and out of the chloroplast. After passing through the TOC complex in the outer chloroplast membrane, polypeptides must pass through the TIC complex ( t ranslocon on the i nner c hloroplast membrane) which is located in the inner chloroplast membrane.
In photosynthesis, the cytochrome b 6 f complex functions to mediate the transfer of electrons and of energy between the two photosynthetic reaction center complexes, Photosystem II and Photosystem I, while transferring protons from the chloroplast stroma across the thylakoid membrane into the lumen. [2]
Chloroplasts can have from 10 to 100 grana. Grana are connected by stroma thylakoids, also called intergranal thylakoids or lamellae. Grana thylakoids and stroma thylakoids can be distinguished by their different protein composition. Grana contribute to chloroplasts' large surface area to volume ratio.
The chloroplast stroma varies from being slightly acidic in the dark to more alkaline in the light. [10] Therefore, in the dark, more FNRs would be recruited and bound to the thylakoid membrane, and in the light, more FNRs would dissociate from the membrane and be free in the stroma.
The structure and function of cytochrome b 6 f (in chloroplasts) is very similar to cytochrome bc 1 (Complex III in mitochondria). Both are transmembrane structures that remove electrons from a mobile, lipid-soluble electron carrier (plastoquinone in chloroplasts; ubiquinone in mitochondria) and transfer them to a mobile, water-soluble electron ...
This creates a H + gradient, making H + ions flow back into the stroma of the chloroplast, providing the energy for the (re)generation of ATP. [citation needed] The photosystem II complex replaced its lost electrons from H 2 O, so electrons are not returned to photosystem II as they would in the analogous cyclic pathway. Instead, they are ...