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Stroma, in botany, refers to the colorless fluid surrounding the grana within the chloroplast. [1] Within the stroma are grana (stacks of thylakoid), the sub-organelles where photosynthesis is started [2] before the chemical changes are completed in the stroma. [3] Photosynthesis occurs in two stages.
Chloroplasts can have from 10 to 100 grana. Grana are connected by stroma thylakoids, also called intergranal thylakoids or lamellae. Grana thylakoids and stroma thylakoids can be distinguished by their different protein composition. Grana contribute to chloroplasts' large surface area to volume ratio.
The inner chloroplast membrane borders the stroma and regulates passage of materials in and out of the chloroplast. After passing through the TOC complex in the outer chloroplast membrane, polypeptides must pass through the TIC complex ( t ranslocon on the i nner c hloroplast membrane) which is located in the inner chloroplast membrane.
The N-terminus of the chlorophyll a-b binding protein extends into the stroma where it is involved with adhesion of granal membranes and photo-regulated by reversible phosphorylation of its threonine residues. [2] Both these processes are believed to mediate the distribution of excitation energy between photosystems I and II.
Within the envelope membranes, in the region called the stroma, there is a system of interconnecting flattened membrane compartments, called the thylakoids.The thylakoid membrane is quite similar in lipid composition to the inner envelope membrane, containing 78% galactolipids, 15.5% phospholipids and 6.5% sulfolipids in spinach chloroplasts. [3]
Cornelis Van Niel proposed in 1931 that photosynthesis is a case of general mechanism where a photon of light is used to photo decompose a hydrogen donor and the hydrogen being used to reduce CO 2. [11] Then in 1939, Robin Hill demonstrated that isolated chloroplasts would make oxygen, but not fix CO
Oxygenic photosynthesis can be performed by plants and cyanobacteria; cyanobacteria are believed to be the progenitors of the photosystem-containing chloroplasts of eukaryotes. Photosynthetic bacteria that cannot produce oxygen have only one photosystem, which is similar to either PSI or PSII .
Photosystem II is present on the thylakoid membranes inside chloroplasts, the site of photosynthesis in green plants. [9] The structure of Photosystem II is remarkably similar to the bacterial reaction center, and it is theorized that they share a common ancestor. The core of Photosystem II consists of two subunits referred to as D1 and D2 ...
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