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Outside the stele lies the endodermis, which is the innermost cell layer of the cortex. The concept of the stele was developed in the late 19th century by French botanists P. E. L. van Tieghem and H. Doultion as a model for understanding the relationship between the shoot and root, and for discussing the evolution of vascular plant morphology. [2]
The endodermis is the boundary between the cortex and the stele. In many seedless plants, such as ferns, the endodermis is a distinct layer of cells immediately outside the vascular cylinder (stele) in roots and shoots. In most seed plants, especially woody types, the endodermis is present in roots but not in stems.
The pericycle is a cylinder of parenchyma or sclerenchyma cells that lies just inside the endodermis and is the outer most part of the stele of plants. [ citation needed ] Although it is composed of non-vascular parenchyma cells, it's still considered part of the vascular cylinder because it arises from the procambium as do the vascular tissues ...
In botany, a cortex is an outer layer of a stem or root in a vascular plant, lying below the epidermis but outside of the vascular bundles. [1] The cortex is composed mostly of large thin-walled parenchyma cells of the ground tissue system and shows little to no structural differentiation. [ 2 ]
The discovery of the Casparian strip dates back to the mid-19th century, and advances in the understanding of the endodermis of plant roots. [15] In 1865, the German botanist Robert Caspary first described the endodermis of the root of plants, found that its cell wall was thickened, and named it Schuchtzscheide.
As the plant grows, new vascular tissue differentiates in the growing tips of the plant. The new tissue is aligned with existing vascular tissue, maintaining its connection throughout the plant. The vascular tissue in plants is arranged in long, discrete strands called vascular bundles. These bundles include both xylem and phloem, as well as ...
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Roots are specialized for the uptake of water, nutrients (including ions for proper function). [12] Similar to the endodermis, the exodermis contains very compact cells and is surrounded by a Casparian band, two features which are used to restrict the flow of water to a symplastic fashion (through the cytoplasm) rather than apoplastic fashion which (through the cell wall) flow through passages ...