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Outside the stele lies the endodermis, which is the innermost cell layer of the cortex. The concept of the stele was developed in the late 19th century by French botanists P. E. L. van Tieghem and H. Doultion as a model for understanding the relationship between the shoot and root, and for discussing the evolution of vascular plant morphology. [2]
The endodermis is the boundary between the cortex and the stele. In many seedless plants, such as ferns, the endodermis is a distinct layer of cells immediately outside the vascular cylinder (stele) in roots and shoots. In most seed plants, especially woody types, the endodermis is present in roots but not in stems.
In botany, a cortex is an outer layer of a stem or root in a vascular plant, lying below the epidermis but outside of the vascular bundles. [1] The cortex is composed mostly of large thin-walled parenchyma cells of the ground tissue system and shows little to no structural differentiation. [ 2 ]
The pericycle is a cylinder of parenchyma or sclerenchyma cells that lies just inside the endodermis and is the outer most part of the stele of plants. [ citation needed ] Although it is composed of non-vascular parenchyma cells, it's still considered part of the vascular cylinder because it arises from the procambium as do the vascular tissues ...
The discovery of the Casparian strip dates back to the mid-19th century, and advances in the understanding of the endodermis of plant roots. [15] In 1865, the German botanist Robert Caspary first described the endodermis of the root of plants, found that its cell wall was thickened, and named it Schuchtzscheide.
In most plants, stems are located above the soil surface, but some plants have underground stems. Stems have several main functions: [3] Support for and the elevation of leaves, flowers, and fruits. The stems keep the leaves in the light and provide a place for the plant to keep its flowers and fruits.
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Parenchyma is a versatile ground tissue that generally constitutes the "filler" tissue in soft parts of plants. It forms, among other things, the cortex (outer region) and pith (central region) of stems, the cortex of roots, the mesophyll of leaves, the pulp of fruits, and the endosperm of seeds.