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In a vascular plant, the stele is the central part of the root or stem [1] containing the tissues derived from the procambium. These include vascular tissue, in some cases ground tissue and a pericycle, which, if present, defines the outermost boundary of the stele.
Polypodiales are unique in bearing sporangia with a vertical annulus interrupted by the stalk and stomium. [2] These sporangial characters were used by Johann Jakob Bernhardi to define a group of ferns he called the "Cathetogyratae"; [3] the Pteridophyte Phylogeny Group has suggested reviving this name as the informal term cathetogyrates, to replace the ambiguously circumscribed term "polypods ...
In 2016, the Pteridophyte Phylogeny Group divided order Polypodiales into six suborders. Pteridaceae is the sole family in suborder Pteridiineae, with 52 genera. The suborder has the same circumscription as Smith et al. used for the family. The phylogenetic relationship between these six suborders is shown in this cladogram: [2]
The Pteridophyte Phylogeny Group classification of 2016 (PPG I) recognized 13 genera. [5] Shortly afterwards, the genus Gastoniella was created for three species formerly placed in Anogramma which a molecular phylogenetic analysis had shown to be distinct.
The forking of the bundle system starts near the base of a terminal cone, which remains erect, of the sympodial development in certain branches. The last method is when "adventitious buds appear in the cortical tissue closely connected with the stelar system of the trunk, and these buds continue their development like typical stems".
The consensus classification produced by the Pteridophyte Phylogeny Group classification in 2016 (PPG I) places all extant (living) lycophytes in the class Lycopodiopsida. [11] There are around 1,290 to 1,340 such species. [12] [13] [11] For more information on the classification of extant lycophytes, see Lycopodiopsida § Classification.
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In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), the family is treated as the subfamily Vittarioideae] of the family Pteridaceae. [ 1 ] The following diagram shows a likely phylogenetic relationship between the Vittarioideae and other subfamilies of the Pteridaceae.