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Pyramidal cells, or pyramidal neurons, are a type of multipolar neuron found in areas of the brain including the cerebral cortex, the hippocampus, and the amygdala. Pyramidal cells are the primary excitation units of the mammalian prefrontal cortex and the corticospinal tract .
There is also a distinct pathway from layer 3 of the EC directly to CA1, often referred to as the temporoammonic or TA-CA1 pathway. Granule cells of the DG send their axons (called "mossy fibers") to CA3. Pyramidal cells of CA3 send their axons to CA1. Pyramidal cells of CA1 send their axons to the subiculum and deep layers of the EC.
The pyramidal tracts include both the corticobulbar tract and the corticospinal tract.These are aggregations of efferent nerve fibers from the upper motor neurons that travel from the cerebral cortex and terminate either in the brainstem (corticobulbar) or spinal cord (corticospinal) and are involved in the control of motor functions of the body.
These are the corticobulbar and corticospinal fibers that make up the pyramidal tracts. About 90% of these fibers leave the pyramids in successive bundles and decussate (cross over) in the anterior median fissure of the medulla oblongata as the pyramidal decussation or motor decussation.
The pyramidal cells in CA3 have a unique type of dendritic spine called a thorny excrescence or thorn, only found in CA3 pyramidal cells and hilar mossy cells. The thorn has a thin single spine with a number of heads. Clusters of thorns sit on a dendrite on a broad stem.
It is the tract of the mossy fiber projections, both inhibitory and excitatory from the granule cells of the dentate gyrus. One mossy fiber may make up to 37 connections to a single pyramidal cell, and innervate around 12 pyramidal cells on top of that. Any given pyramidal cell in the stratum lucidum may get input from as many as 50 granule cells.
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A human neocortical pyramidal cell that has been stained using Golgi's method. The cell is named after its characteristic triangular-shaped soma. Since there is a broad range of functions performed by different types of neurons in diverse parts of the nervous system, there is a wide variety in the size, shape, and electrochemical properties of ...