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In most modern reptiles and amphibians, the quadratojugal is a prominent, straplike bone in the skull and provides structural integrity in the postorbital region of the skull. [6] In many reptiles, the inner face of the quadratojugal also connects to the quadrate bone which forms the cranium's contribution to the jaw joint.
The jugal bone is located on either side of the skull in the circumorbital region. It is the origin of several masticatory muscles in the skull. [1] The jugal and lacrimal bones are the only two remaining from the ancestral circumorbital series: the prefrontal, postfrontal, postorbital, jugal, and lacrimal bones.
The quadrate bone is a skull bone in most tetrapods, including amphibians, sauropsids (reptiles, birds), and early synapsids. In most tetrapods, the quadrate bone connects to the quadratojugal and squamosal bones in the skull, and forms upper part of the jaw joint.
The jugal, or cheek-bone, is a skull bone that defines the lower border of the orbit and connects to the maxilla anteriorly and to the quadratojugal posteriorly. Its posterior end is bifurcated at its articulation with the quadratojugal, which is considered a synapomorphy of dinosaurs. [1]: 140 [25]: 12
The jugal and quadratojugal have pinched tubercular ornamentation on the lateral sides. [1] The quadratojugal has the elongated and slender shape distinct to the Varanopids, as well as a slender groove on the dorsal edge. This serves as an attachment for the squamosal, which extends anteriorly under the lateral temporal fenestra. [1]
The squamosal is a skull bone found in most reptiles, amphibians, and birds. In fishes, it is also called the pterotic bone. [1] In most tetrapods, the squamosal and quadratojugal bones form the cheek series of the skull. [2] The bone forms an ancestral component of the dermal roof and is typically thin compared to other skull bones. [3]
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The skull of Tanycolagreus is less well known than its postcranial anatomy, and only the following elements have been found: left nasal, left lacrimal, left premaxilla and one premaxillary tooth, left postorbital, left quadratojugal, incomplete left squamosal, right quadrate, right splenial, left articular, and two