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The male gametophyte will develop via one or two rounds of mitosis inside the anther. This creates a 2 or 3 celled male gametophyte which becomes known as the pollen grain once dehiscing occurs. [18] One cell is the tube cell, and the remaining cell/cells are the sperm cells. [19]
It is the biological process of gametogenesis during which cells that are haploid or diploid divide to create other cells. It can take place either through mitotic or meiotic division of diploid gametocytes into different cells depending on an organism's biological life cycle. For instance, gametophytes in plants undergo mitosis to produce gametes.
Sperm cells are derived by mitosis of the generative cell during pollen tube elongation. The vegetative cell is responsible for pollen tube development. Double-strand breaks in DNA that arise appear to be efficiently repaired in the generative cell, but not in the vegetative cell, during the transport process to the female gametophyte. [32]
However, the parent sporophyte may be monoecious, producing both male and female gametophytes or dioecious, producing gametophytes of one gender only. Seed plant gametophytes are extremely reduced in size; the archegonium consists only of a small number of cells, and the entire male gametophyte may be represented by only two cells. [27]
The different stages of mitosis altogether define the mitotic phase (M phase) of a cell cycle—the division of the mother cell into two daughter cells genetically identical to each other. [ 3 ] The process of mitosis is divided into stages corresponding to the completion of one set of activities and the start of the next.
These microspore mother cells, also called microsporocytes, then undergo meiosis and become four microspore haploid cells. These new microspore cells then undergo mitosis and form a tube cell and a generative cell. The generative cell then undergoes mitosis one more time to form two male gametes, also called sperm.
The eukaryotic cell cycle consists of four distinct phases: G 1 phase, S phase (synthesis), G 2 phase (collectively known as interphase) and M phase (mitosis and cytokinesis). M phase is itself composed of two tightly coupled processes: mitosis, in which the cell's nucleus divides, and cytokinesis, in which the cell's cytoplasm and cell membrane divides forming two daughter cells.
These spores then germinate and divide by mitosis to form a haploid multicellular phase, the gametophyte, which produces gametes directly by mitosis. This type of life cycle, involving alternation between two multicellular phases, the sexual haploid gametophyte and asexual diploid sporophyte, is known as alternation of generations.