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The calcichordate hypothesis, formulated by British Museum paleontologist Richard Jefferies, holds that each separate lineage of chordate (Cephalochordates, Urochordates, Craniates) evolved from its own lineage of mitrate, and thus the echinoderms and the chordates are sister groups, with the hemichordates as an out-group. [1]
Spotted gar, Lepisosteus oculatus This article contains a list of all of the classes and orders that are located in the Phylum Chordata.. The subphyla Tunicata and Vertebrata are in the unranked Olfactores clade, while the subphylum Cephalochordata is not.
Hemichordates ("half chordates") have some features similar to those of chordates: branchial openings that open into the pharynx and look rather like gill slits; stomochords, similar in composition to notochords, but running in a circle round the "collar", which is ahead of the mouth; and a dorsal nerve cord—but also a smaller ventral nerve cord.
The Cambrian chordates are characterised by the presence of segmented muscle blocks called myomeres and notochord, the two defining features of chordates. Before the full understanding of Cambrian fossils, chordates as members the most advanced phylum were believed to appear on Earth much later than the Cambrian.
Some biologists have proposed that the Hemichordates (specifically the Enteropneusta) may represent an intermediate body plan in the evolution of the "inverted" state of the chordates. [10] Though they are considered deuterostomes, the dorsoventral axis of hemichordates retains features of both protostomes and chordates.
The notochord is the defining feature (synapomorphy) of chordates, and was present throughout life in many of the earliest chordates. Although the stomochord of hemichordates was once thought to be homologous or from a common lineal origin, it is now viewed as analogous, convergent , or from a different lineal origin. [ 22 ]
Pikaia gracilens is an extinct, primitive chordate marine animal known from the Middle Cambrian Burgess Shale of British Columbia.Described in 1911 by Charles Doolittle Walcott as an annelid, and in 1979 by Harry B. Whittington and Simon Conway Morris as a chordate, it became "the most famous early chordate fossil", [1] or "famously known as the earliest described Cambrian chordate". [2]
A craniate is a member of the Craniata (sometimes called the Craniota), a proposed clade of chordate animals with a skull of hard bone or cartilage.Living representatives are the Myxini (hagfishes), Hyperoartia (including lampreys), and the much more numerous Gnathostomata (jawed vertebrates).