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Psilotum is a genus of fern-like vascular plants.It is one of two genera in the family Psilotaceae commonly known as whisk ferns, the other being Tmesipteris.Plants in these two genera were once thought to be descended from the earliest surviving vascular plants, but more recent phylogenies place them as basal ferns, as a sister group to Ophioglossales.
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Higher up it became elliptical in cross-section, corresponding to the two rows of fertile branching units. Within the branches of these units, the strand was more-or-less rectangular. The conducting elements of the xylem, the tracheids , were of the so-called 'P-type' in which the walls were strengthened by ladder-like (scalariform) bars with ...
The stem anatomy of ferns is more complicated than that of dicots because fern stems often have one or more leaf gaps in cross section. A leaf gap is where the vascular tissue branches off to a frond. In cross section, the vascular tissue does not form a complete cylinder where a leaf gap occurs.
It forms a protective covering on the leaf vein and consists of one or more cell layers, usually parenchyma. Loosely-arranged mesophyll cells lie between the bundle sheath and the leaf surface. The Calvin cycle is confined to the chloroplasts of these bundle sheath cells in C 4 plants. C 2 plants also use a variation of this structure. [1]
An interesting case is that of Psilotum, which has a (simple) protostele, and enations devoid of vascular tissue. Some species of Psilotum have a single vascular trace that terminates at the base of the enations. [2] Consequently, Psilotum was long thought to be a "living fossil" closely related to early land plants (rhyniophytes).
Psilotum has been interpreted as producing sporangia (fused in a synangium) on the terminus of a stem. Equisetum always produce strobili, but the structures bearing sporangia (sporangiophores) have been interpreted as modified stems. The sporangia, despite being recurved are interpreted as terminal.
The Novak–Tyson Model is a non-linear dynamics framework developed in the context of cell-cycle control by Bela Novak and John J. Tyson. It is a prevalent theoretical model that describes a hysteretic , bistable bifurcation of which many biological systems have been shown to express.