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The jugal bone is located on either side of the skull in the circumorbital region. It is the origin of several masticatory muscles in the skull. [1] The jugal and lacrimal bones are the only two remaining from the ancestral circumorbital series: the prefrontal, postfrontal, postorbital, jugal, and lacrimal bones. [2]
In tetrapods with a quadratojugal bone, it often forms a portion of the jaw joint. Developmentally, the quadratojugal bone is a dermal bone in the temporal series, forming the original braincase. The squamosal and quadratojugal bones together form the cheek region [4] and may provide muscular attachments for facial muscles. [5]
These extended margins of thinned bone are called supratemporal fossae. Synapsids, including mammals, have one temporal fenestra, which is ventrally bordered by a zygomatic arch composed of the jugal and squamosal bones. This single temporal fenestra is homologous to the infratemporal fenestra, as displayed most clearly by early synapsids. [2]
It is composed of 270 bones at the time of birth, [2] but later decreases to 206: 80 bones in the axial skeleton and 126 bones in the appendicular skeleton. 172 of 206 bones are part of a pair and the remaining 34 are unpaired. [3] Many small accessory bones, such as sesamoid bones, are not included in this.
The epijugal is a dermal ossification unique to ceratopsians. It caps the bottom end of the downward-facing, triangular flange of the jugal that is typical for the group. In Arrhinoceratops and Pentaceratops, the epijugal forms a distinct jugal horn. The bone can be found in Yamaceratops and all ceratopsians more derived than the latter.
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The axial skeleton (80 bones) is formed by the vertebral column (32–34 bones; the number of the vertebrae differs from human to human as the lower 2 parts, sacral and coccygeal bone may vary in length), a part of the rib cage (12 pairs of ribs and the sternum), and the skull (22 bones and 7 associated bones).