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In trees, the phloem is the innermost layer of the bark, hence the name, derived from the Ancient Greek word φλοιός (phloiós), meaning "bark". [3] [4] The term was introduced by Carl Nägeli in 1858. [5] [6] Different types of phloem can be distinguished. The early phloem formed in the growth apices is called protophloem.
Cross section of celery stalk, showing vascular bundles, which include both phloem and xylem Detail of the vasculature of a bramble leaf Translocation in vascular plants. Vascular tissue is a complex transporting tissue, formed of more than one cell type, found in vascular plants. The primary components of vascular tissue are the xylem and ...
Xylem cells are dead, hard-walled hollow cells arranged to form files of tubes that function in water transport. A tracheid cell wall usually contains the polymer lignin. [citation needed] The phloem, on the other hand, consists of living cells called sieve-tube members. Between the sieve-tube members are sieve plates, which have pores to allow ...
Pressure flow hypothesis: Sugars produced in the leaves and other green tissues are kept in the phloem system, creating a solute pressure differential versus the xylem system carrying a far lower load of solutes—water and minerals. The phloem pressure can rise to several MPa, [12] far higher than atmospheric pressure. Selective inter ...
Another important component of phloem sap is nitrogen. Nitrogen is usually not being transported in its ionic form. [11] Instead it is incorporated in amino acids such as glutamate and aspartate. [11] Hormones, inorganic ions, RNA, and proteins are found in the phloem sap as well. [11] [12] Xylem sap is mostly made of water.
In old stems the epidermal layer, cortex, and primary phloem become separated from the inner tissues by thicker formations of cork. Due to the thickening cork layer these cells die because they do not receive water and nutrients. This dead layer is the rough corky bark that forms around tree trunks and other stems.
Primarily, phloem carries dissolved food substances throughout the plant. This conduction system is composed of sieve-tube member and companion cells, that are without secondary walls. The parent cells of the vascular cambium produce both xylem and phloem. This usually also includes fibers, parenchyma and ray cells.
The transport itself happens in the stem, which exists in two forms: xylem and phloem. Both these tissues are present in a vascular bundle, which in addition will include supporting and protective tissues. There is also a tissue between xylem and phloem, which is the cambium.