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The activity of plasmodesmata are linked to physiological and developmental processes within plants. There is a hormone signaling pathway that relays primary cellular signals via the plasmodesmata. There are also patterns of environmental, physiological, and developmental cues that show relation to plasmodesmata function.
A desmotubule is an endomembrane derived structure of the plasmodesmata that connects the endoplasmic reticulum of two adjacent plant cells. [1] [2] The desmotubule is not actually a tubule but a compact, cylindrical segment of the ER that is found within the larger tubule structure of the plasmodesmata pore. [3]
All of the cellular functions of a sieve-tube element are carried out by the (much smaller) companion cell, a typical nucleate plant cell except the companion cell usually has a larger number of ribosomes and mitochondria. The dense cytoplasm of a companion cell is connected to the sieve-tube element by plasmodesmata. [10]
Passive phloem loading transports solutes freely through plasmodesma in the symplast of the minor veins of leaves. Active transport occurs apoplastically and does not use plasmodesmata. An intermediate type of loading exists that uses symplastic transport but utilizes a size-exclusion mechanism to ensure diffusion is a one-way process between ...
The apoplastic and symplastic pathways. The symplast of a plant is the region enclosed by the cell membranes, within which water and solutes can diffuse freely. By contrast the apoplast is any fluid-filled space within the cell wall and extracellular space. [1]
Plant cells are surrounded by solid cell walls, therefore transport through plasmodesmata is the preferred path for virions to move between plant cells. Plants have specialized mechanisms for transporting mRNAs through plasmodesmata, and these mechanisms are thought to be used by RNA viruses to spread from one cell to another. [2]
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The sucrose then diffuses through the plasmodesmata from the companion cells to the sieve tube elements. As a result, the concentration of sucrose increases in the sieve tube elements. This causes water to move into the sieve tube element by osmosis, creating pressure that pushes the sap down the tube.