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The Bak–Sneppen model is a simple model of co-evolution between interacting species. It was developed to show how self-organized criticality may explain key features of the fossil record, such as the distribution of sizes of extinction events and the phenomenon of punctuated equilibrium. It is named after Per Bak and Kim Sneppen.
However, the punctuational equilibrium model may still be inferred from both the observation of stasis and examples of rapid and episodic speciation events documented in the fossil record. [70] Dawkins also emphasizes that punctuated equilibrium has been "oversold by some journalists", [71] but partly due to Eldredge and Gould's "later writings ...
Punctuated gradualism is considered to be a variation of these models, lying somewhere in between the phyletic gradualism model and the punctuated equilibrium model. It states that speciation is not needed for a lineage to rapidly evolve from one equilibrium to another but may show rapid transitions between long-stable states.
Dawkins could, should, and probably would accept it; in The Ancestor's Tale, he has an inclusive view of speciation mechanisms." (p. 100) Thus, while "Gould somewhat overstates the adherence of orthodoxy to strict extrapolationism", punctuated equilibrium is more important than some of the more "ungenerous treatment" that has been meted out.
Gould's most significant contribution to evolutionary biology was the theory of punctuated equilibrium [2] developed with Niles Eldredge in 1972. [3] The theory proposes that most evolution is characterized by long periods of evolutionary stability, infrequently punctuated by swift periods of branching speciation .
Gould's initial work on punctuated equilibrium was coauthored with Niles Eldredge. [20] Gould's model of punctuated equilibrium drew attention to episodic bursts of evolutionary change followed by periods of morphological stability. He challenged the conventional model of gradual, continuous change - called phyletic gradualism. [21]
Next, thrust in an inward and upward motion on the diaphragm. This will force air out of the lungs and remove the blockage. Repeat these abdominal thrusts up to five times, the doctor advised.
Punctuated equilibrium [ edit ] In the 1980s, the American palaeontologists Stephen Jay Gould and Niles Eldredge argued for an extended synthesis based on their idea of punctuated equilibrium , the role of species selection shaping large scale evolutionary patterns and natural selection working on multiple levels extending from genes to species.