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However, most tetrapod species today are amniotes, most of which are terrestrial tetrapods whose branch evolved from earlier tetrapods early in the Late Carboniferous. The key innovation in amniotes over amphibians is the amnion, which enables the eggs to retain their aqueous contents on land, rather than needing to stay in water.
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The evolution of tetrapods began about 400 million years ago in the Devonian Period with the earliest tetrapods evolved from lobe-finned fishes. [1] Tetrapods (under the apomorphy-based definition used on this page) are categorized as animals in the biological superclass Tetrapoda, which includes all living and extinct amphibians, reptiles, birds, and mammals.
Tetrapodomorpha (also known as Choanata [3]) is a clade of vertebrates consisting of tetrapods (four-limbed vertebrates) and their closest sarcopterygian relatives that are more closely related to living tetrapods than to living lungfish.
Both its major subgroups are successful today: Actinopterygii includes most extant bony fish species, and Sarcopterygii includes the tetrapods. Euteleostomes originally all had an endochondral bone, fins with lepidotrichs (fin rays), jaws lined by maxillary, premaxillary, and dentary bones composed of dermal bone, and lungs.
Amphibians today generally remain semi-aquatic, living the first stage of their lives as fish-like tadpoles. Several groups of tetrapods, such as the reptillian snakes and mammalian cetaceans, have lost some or all of their limbs, and many tetrapods have returned to partially aquatic or (in the case of cetaceans and sirenians) fully aquatic lives.
Traditionally, Ichthyostega was considered part of an order named for it, the "Ichthyostegalia". However, this group represents a paraphyletic grade of primitive stem-tetrapods and is not used by many modern researchers. Phylogenetic analysis has shown Ichthyostega is intermediate between other primitive stegocephalian stem-tetrapods. The ...
Tiktaalik, a tetrapodomorph with wrists, straddles the fish-tetrapod divide. The Stem Tetrapoda are a cladistically defined group, consisting of all animals more closely related to extant four-legged vertebrates than to their closest extant relatives (the lungfish), but excluding the crown group Tetrapoda.