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In trees, the phloem is the innermost layer of the bark, hence the name, derived from the Ancient Greek word φλοιός (phloiós), meaning "bark". [3] [4] The term was introduced by Carl Nägeli in 1858. [5] [6] Different types of phloem can be distinguished. The early phloem formed in the growth apices is called protophloem.
The phloem is the living portion of the vascular system of a plant, and serves to move sugars and photosynthate from source cells to sink cells. Phloem tissue is made of sieve elements and companion cells, and is surrounded by parenchyma cells. The sieve element cells work as the main player in transport of phloem sap.
Parenchyma is a versatile ground tissue that generally constitutes the "filler" tissue in soft parts of plants. It forms, among other things, the cortex (outer region) and pith (central region) of stems, the cortex of roots, the mesophyll of leaves, the pulp of fruits, and the endosperm of seeds.
Lateral meristems, the form of secondary plant growth, add growth to the plants in their diameter. This is primarily observed in perennial dicots that survive from year to year. There are two types of lateral meristems: vascular cambium and cork cambium. In vascular cambium, the primary phloem and xylem are produced by the apical meristem.
Two kinds of vascular tissue occur in plants: xylem and phloem. Phloem and xylem are closely associated with one another and are typically located immediately adjacent to each other in the plant. The combination of one xylem and one phloem strand adjacent to each other is known as a vascular bundle. [14]
Xylem is one of the two types of transport tissue in vascular plants, the other being phloem; both of these are part of the vascular bundle. The basic function of the xylem is to transport water upward from the roots to parts of the plants such as stems and leaves, but it also transports nutrients .
Sieve cells are long, conducting cells in the phloem that do not form sieve tubes. The major difference between sieve cells and sieve tube members is the lack of sieve plates in sieve cells. [ 1 ] They have a very narrow diameter and tend to be longer in length than sieve tube elements as they are generally associated with albuminous cells. [ 4 ]
The fascicular and interfascicular cambia thus join up to form a ring (in three dimensions, a tube) which separates the primary xylem and primary phloem, the cambium ring. The vascular cambium produces secondary xylem on the inside of the ring, and secondary phloem on the outside, pushing the primary xylem and phloem apart.