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In trees, the phloem is the innermost layer of the bark, hence the name, derived from the Ancient Greek word φλοιός (phloiós), meaning "bark". [3] [4] The term was introduced by Carl Nägeli in 1858. [5] [6] Different types of phloem can be distinguished. The early phloem formed in the growth apices is called protophloem.
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The first xylem to develop is called 'protoxylem'. In appearance, protoxylem is usually distinguished by narrower vessels formed of smaller cells. Some of these cells have walls that contain thickenings in the form of rings or helices. Functionally, protoxylem can extend: the cells can grow in size and develop while a stem or root is elongating.
Primarily, phloem carries dissolved food substances throughout the plant. This conduction system is composed of sieve-tube member and companion cells, that are without secondary walls. The parent cells of the vascular cambium produce both xylem and phloem. This usually also includes fibers, parenchyma and ray cells.
Glands secreting a sugary liquid—the nectar—in flowers pollinated by insects are called nectaries. Nectaries may occur on the floral stalk or on any floral organ: sepal, petal, stamen, or ovary. The hydathode structures discharge water—a phenomenon called guttation through openings in margins or tips of leaves.
According to the hypothesis, the high concentration of organic substances, particularly sugar, inside the phloem at a source such as a leaf creates a diffusion gradient (osmotic gradient) that draws water into the cells from the adjacent xylem. This creates turgor pressure, also called hydrostatic pressure, in the phloem. The hypothesis states ...
Phloem was introduced by Carl Nägeli in 1858 after the discovery of sieve elements. Since then, multiple studies have been conducted on how sieve elements function in phloem in terms of working as a transport mechanism. [2] An example of analysis of phloem through sieve elements was conducted in the study of Arabidopsis leaves.
An endodermis generally surrounds the stele. A centrarch (protoxylem in the center of a metaxylem cylinder) haplostele is prevalent in members of the rhyniophyte grade, such as Rhynia. [6] actinostele – a variation of the protostele in which the core is lobed or fluted. [7] This stele is found in many species of club moss (Lycopodium and ...