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C 4 carbon fixation or the Hatch–Slack pathway is one of three known photosynthetic processes of carbon fixation in plants. It owes the names to the 1960s discovery by Marshall Davidson Hatch and Charles Roger Slack. [1] C 4 fixation is an addition to the ancestral and more common C 3 carbon fixation.
Maize (Zea mays, Poaceae) is the most widely cultivated C 4 plant.[1]In botany, C 4 carbon fixation is one of three known methods of photosynthesis used by plants. C 4 plants increase their photosynthetic efficiency by reducing or suppressing photorespiration, which mainly occurs under low atmospheric CO 2 concentration, high light, high temperature, drought, and salinity.
As a plant that uses C 4 carbon fixation, maize is a considerably more water-efficient crop than plants that use C 3 carbon fixation such as alfalfa and soybeans. Maize is most sensitive to drought at the time of silk emergence, when the flowers are ready for pollination.
Phosphoenolpyruvate carboxylase (also known as PEP carboxylase, PEPCase, or PEPC; EC 4.1.1.31, PDB ID: 3ZGE) is an enzyme in the family of carboxy-lyases found in plants and some bacteria that catalyzes the addition of bicarbonate (HCO 3 −) to phosphoenolpyruvate (PEP) to form the four-carbon compound oxaloacetate and inorganic phosphate: [1]
Through photosynthesis, plants use CO 2 from the atmosphere, water from the ground, and energy from the sun to create sugars used for growth and fuel. [22] While using these sugars as fuel releases carbon back into the atmosphere (photorespiration), growth stores carbon in the physical structures of the plant (i.e. leaves, wood, or non-woody stems). [23]
Biological carbon fixation, or сarbon assimilation, is the process by which living organisms convert inorganic carbon (particularly carbon dioxide) to organic compounds. These organic compounds are then used to store energy and as structures for other biomolecules .
Carbon on Earth naturally occurs in two stable isotopes, with 98.9% in the form of 12 C and 1.1% in 13 C. [1] [8] The ratio between these isotopes varies in biological organisms due to metabolic processes that selectively use one carbon isotope over the other, or "fractionate" carbon through kinetic or thermodynamic effects. [1]
The fixation or reduction of carbon dioxide is a process in which carbon dioxide combines with a five-carbon sugar, ribulose 1,5-bisphosphate, to yield two molecules of a three-carbon compound, glycerate 3-phosphate, also known as 3-phosphoglycerate.