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The iris consists of two layers: the front pigmented fibrovascular layer known as a stroma and, behind the stroma, pigmented epithelial cells.. The stroma is connected to a sphincter muscle (sphincter pupillae), which contracts the pupil in a circular motion, and a set of dilator muscles (dilator pupillae), which pull the iris radially to enlarge the pupil, pulling it in folds.
The pupil constricts when the iris sphincter muscle is stimulated and contracts. In humans, it functions to constrict the pupil in bright light (pupillary light reflex) or during accommodation. [citation needed] In lower animals [clarification needed], the muscle cells themselves are photosensitive causing iris action without brain input. [5]
The seventh muscle is the levator palpebrae superioris muscle. When the muscles exert different tensions, a torque is exerted on the globe that causes it to turn, in almost pure rotation, with only about one millimeter of translation. [8] Thus, the eye can be considered as undergoing rotations about a single point in the centre of the eye.
Muscles around the iris change the size of the pupil, regulating the amount of light that enters the eye [3] and reducing aberrations when there is enough light. [4] The eyes of most cephalopods , fish , amphibians and snakes have fixed lens shapes, and focusing is achieved by telescoping the lens in a similar manner to that of a camera .
The iris dilator muscle (pupil dilator muscle, pupillary dilator, radial muscle of iris, radiating fibers), is a smooth muscle [2] of the eye, running radially in the iris and therefore fit as a dilator. The pupillary dilator consists of a spokelike arrangement of modified contractile cells called myoepithelial cells.
Structure of the iris and surrounding parts showing the stroma of iris (stroma iridis). The stroma connects to a sphincter muscle ( sphincter pupillae ), which contracts the pupil in a circular motion, and a set of dilator muscles ( dilator pupillae ) which pull the iris radially to enlarge the pupil, pulling it in folds.
This double membrane is often considered continuous with the retina and a rudiment of the embryological correspondent to the retina. The inner layer is unpigmented until it reaches the iris, where it takes on pigment. The retina ends at the ora serrata. The space between the ciliary body and the base of the iris is the ciliary sulcus. [4]
Neuroepithelium forms the retina, ciliary body, iris, and optic nerves. Surface ectoderm forms the lens, corneal epithelium and eyelid. The extracellular mesenchyme forms the sclera, the corneal endothelium and stroma, blood vessels, muscles, and vitreous.