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The evolution of tetrapods began about 400 million years ago in the Devonian Period with the earliest tetrapods evolved from lobe-finned fishes. [1] Tetrapods (under the apomorphy-based definition used on this page) are categorized as animals in the biological superclass Tetrapoda, which includes all living and extinct amphibians, reptiles, birds, and mammals.
In effect, "tetrapod" is a name reserved solely for animals which lie among living tetrapods, so-called crown tetrapods. This is a node-based clade , a group with a common ancestry descended from a single "node" (the node being the nearest common ancestor of living species).
The universal tetrapod characteristics of front limbs that bend backward at the elbow and hind limbs that bend forward at the knee can plausibly be traced to early tetrapods living in shallow water. [18] Panderichthys is a 90–130 cm (35–50 in) long fish from the Late Devonian period (380 Mya). It has a large tetrapod-like head.
Early fossil tetrapods have been found in marine sediments, and because fossils of primitive tetrapods in general are found scattered all around the world, they must have spread by following the coastal lines — they could not have lived in freshwater only. Fossil Illuminates Evolution of Limbs from Fins Scientific American, 2 2 April 2004.
Tiktaalik, a lobe-finned fish with some anatomical features similar to early tetrapods. It has been suggested to be a transitional species between fish and tetrapods. [81] 365 Ma Acanthostega is one of the earliest vertebrates capable of walking. [82] 363 Ma By the start of the Carboniferous Period, the Earth begins to resemble its present state.
Nine tetrapod trackways from three sites have been reported from the Valentia Slate Formation of Valentia Island, Ireland. [ 2 ] [ 3 ] The Valentia Slate Formation is composed mostly of purple coloured fine-grained sandstones and siltstones interpreted to represent a fluvial setting.
Stegocephalians include both the modern lineage of limbed vertebrates (the crown group tetrapods, including modern amphibians, reptiles, birds and mammals) as well as a portion of the stem group, the earliest limbed tetrapodomorphs such as Ichthyostega and Acanthostega, which evolved in the Devonian period long before the origin of the crown group.
The fins of lobe-finned fish differ from those of all other fish in that each is borne on a fleshy, lobelike, scaly stalk extending from the body. The pectoral and pelvic fins are articulated in ways resembling the tetrapod limbs they were the precursors to. The fins evolved into the legs of the first tetrapod land vertebrates, amphibians.