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Phloem (/ ˈ f l oʊ. əm /, FLOH-əm) is the living tissue in vascular plants that transports the soluble organic compounds made during photosynthesis and known as photosynthates, in particular the sugar sucrose, [1] to the rest of the plant. This transport process is called translocation. [2]
This early water transport took advantage of the cohesion-tension mechanism inherent in water. Water has a tendency to diffuse to areas that are drier, and this process is accelerated when water can be wicked along a fabric with small spaces. In small passages, such as that between the plant cell walls (or in tracheids), a column of water ...
Cross section of celery stalk, showing vascular bundles, which include both phloem and xylem Detail of the vasculature of a bramble leaf Translocation in vascular plants. Vascular tissue is a complex transporting tissue, formed of more than one cell type, found in vascular plants. The primary components of vascular tissue are the xylem and ...
Monosaccharides, though generally more complex sugars, are made in the "dark" reactions. The term "light" reaction can be confusing as some "dark" reactions require light to be active. [citation needed] Photoassimilate movement through plants from "source to sink" using xylem and phloem is of biological significance.
The phloem sugar is consumed by cellular respiration or converted into starch, which is insoluble and exerts no osmotic effect. With much of the sucrose having been removed, the water exits the phloem by osmosis or is drawn by transpiration into nearby xylem vessels, lowering the turgor pressure within the phloem. [4]
The fascicular and interfascicular cambia thus join up to form a ring (in three dimensions, a tube) which separates the primary xylem and primary phloem, the cambium ring. The vascular cambium produces secondary xylem on the inside of the ring, and secondary phloem on the outside, pushing the primary xylem and phloem apart.
The process for this is retting, and can be performed by micro-organisms either on land (nowadays the most important) or in water, or by chemicals (for instance high pH and chelating agents), or by pectinolytic enzymes. In the phloem, bast fibres occur in bundles that are glued together by pectin and calcium ions. More intense retting separates ...
An endodermis generally surrounds the stele. A centrarch (protoxylem in the center of a metaxylem cylinder) haplostele is prevalent in members of the rhyniophyte grade, such as Rhynia. [6] actinostele – a variation of the protostele in which the core is lobed or fluted. [7] This stele is found in many species of club moss (Lycopodium and ...