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The jugal bone is located on either side of the skull in the circumorbital region. It is the origin of several masticatory muscles in the skull. [1] The jugal and lacrimal bones are the only two remaining from the ancestral circumorbital series: the prefrontal, postfrontal, postorbital, jugal, and lacrimal bones. [2]
In tetrapods with a quadratojugal bone, it often forms a portion of the jaw joint. Developmentally, the quadratojugal bone is a dermal bone in the temporal series, forming the original braincase. The squamosal and quadratojugal bones together form the cheek region [4] and may provide muscular attachments for facial muscles. [5]
The epijugal is a dermal ossification unique to ceratopsians. It caps the bottom end of the downward-facing, triangular flange of the jugal that is typical for the group. In Arrhinoceratops and Pentaceratops, the epijugal forms a distinct jugal horn. The bone can be found in Yamaceratops and all ceratopsians more derived than the latter.
These extended margins of thinned bone are called supratemporal fossae. Synapsids, including mammals, have one temporal fenestra, which is ventrally bordered by a zygomatic arch composed of the jugal and squamosal bones. This single temporal fenestra is homologous to the infratemporal fenestra, as displayed most clearly by early synapsids. [2]
The squamosal is a skull bone found in most reptiles, amphibians, and birds. In fishes, it is also called the pterotic bone. [1] In most tetrapods, the squamosal and quadratojugal bones form the cheek series of the skull. [2] The bone forms an ancestral component of the dermal roof and is typically thin compared to other skull bones. [3]
It is composed of 270 bones at the time of birth, [2] but later decreases to 206: 80 bones in the axial skeleton and 126 bones in the appendicular skeleton. 172 of 206 bones are part of a pair and the remaining 34 are unpaired. [3] Many small accessory bones, such as sesamoid bones, are not included in this.
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Pachycephalosaurs, with their dome-shaped heads, are commonly thought to have used their thick skulls for butting into each other. Their brains and vertebrae are positioned in ways to protect from the impact of head-butting. Also, the columnar structure found in bone remains is consistent with models used to recreate the practice of head ...