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A gene cluster is a group of two or more genes found within an organism's DNA that encode similar polypeptides or proteins which collectively share a generalized function and are often located within a few thousand base pairs of each other.
However, PSII has an additional function over the bacterial system. At the oxidising side of PSII, a redox-active residue in the D1 protein reduces P680, the oxidised tyrosine then withdrawing electrons from a manganese cluster, which in turn withdraw electrons from water, leading to the splitting of water and the formation of molecular oxygen.
Chloroplast DNA (cpDNA), also known as plastid DNA (ptDNA) is the DNA located in chloroplasts, which are photosynthetic organelles located within the cells of some eukaryotic organisms. Chloroplasts, like other types of plastid , contain a genome separate from that in the cell nucleus .
Four different subunits were found to be important for the function of the photosynthetic reaction center. The L and M subunits, shown in blue and purple in the image of the structure, both span the lipid bilayer of the plasma membrane. They are structurally similar to one another, both having 5 transmembrane alpha helices. [6]
The main function of PSII is to efficiently split water into oxygen molecules and protons. PSII will provide a steady stream of electrons to PSI, which will boost these in energy and transfer them to NADP + and H + to make NADPH. The hydrogen from this NADPH can then be used in a number of different processes within the plant. [2]
It is a metallo-oxo cluster comprising four manganese ions (in oxidation states ranging from +3 to +4) [6] and one divalent calcium ion. When it oxidizes water, producing oxygen gas and protons, it sequentially delivers the four electrons from water to a tyrosine (D1-Y161) sidechain and then to P680 itself.
P680 + is the strongest biological oxidizing agent known, with an estimated redox potential of ~1.3 V. [3] This makes it possible to oxidize water during oxygenic photosynthesis. P680 + recovers its lost electron by oxidizing water via the oxygen-evolving complex , which regenerates P680.
The light-harvesting complex (or antenna complex; LH or LHC) is an array of protein and chlorophyll molecules embedded in the thylakoid membrane of plants and cyanobacteria, which transfer light energy to one chlorophyll a molecule at the reaction center of a photosystem.