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English: xylem (blue) carries water from the roots upwards phloem (orange) carries products of photosynthesis from the place of their origin (source) to organs where they are needed (roots, storage organs, flowers, fruits – sink); note that e.g. the storage organs may be source and leaves may be sink at the beginning of the growing season
In trees, the phloem is the innermost layer of the bark, hence the name, derived from the Ancient Greek word φλοιός (phloiós), meaning "bark". [3] [4] The term was introduced by Carl Nägeli in 1858. [5] [6] Different types of phloem can be distinguished. The early phloem formed in the growth apices is called protophloem.
The phloem is the living portion of the vascular system of a plant, and serves to move sugars and photosynthate from source cells to sink cells. Phloem tissue is made of sieve elements and companion cells, and is surrounded by parenchyma cells. The sieve element cells work as the main player in transport of phloem sap.
Pressure flow hypothesis: Sugars produced in the leaves and other green tissues are kept in the phloem system, creating a solute pressure differential versus the xylem system carrying a far lower load of solutes—water and minerals. The phloem pressure can rise to several MPa, [12] far higher than atmospheric pressure. Selective inter ...
The phloem sugar is consumed by cellular respiration or converted into starch, which is insoluble and exerts no osmotic effect. With much of the sucrose having been removed, the water exits the phloem by osmosis or is drawn by transpiration into nearby xylem vessels, lowering the turgor pressure within the phloem. [ 4 ]
The cells in vascular tissue are typically long and slender. Since the xylem and phloem function in the conduction of water, minerals, and nutrients throughout the plant, it is not surprising that their form should be similar to pipes. The individual cells of phloem are connected end-to-end, just as the sections of a pipe might be.
Since the valuable fibres are located in the phloem, they must often be separated from the woody core, the xylem, and sometimes also from the epidermis. The process for this is retting , and can be performed by micro-organisms either on land (nowadays the most important) or in water, or by chemicals (for instance high pH and chelating agents ...
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