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Phloem (/ ˈ f l oʊ. əm /, FLOH-əm) is the living tissue in vascular plants that transports the soluble organic compounds made during photosynthesis and known as photosynthates, in particular the sugar sucrose, [1] to the rest of the plant.
Vascular tissue is a complex conducting tissue, formed of more than one cell type, found in vascular plants. The primary components of vascular tissue are the xylem and phloem. These two tissues transport fluid and nutrients internally. There are also two meristems associated with vascular tissue: the vascular cambium and the cork cambium.
Both these tissues are present in a vascular bundle, which in addition will include supporting and protective tissues. There is also a tissue between xylem and phloem, which is the cambium. The xylem typically lies towards the axis with phloem positioned away from the axis . In a stem or root this means that the xylem is closer to the centre of ...
Xylem is the water-conducting tissue, and the secondary xylem provides the raw material for the forest products industry. [26] Xylem and phloem tissues each play a part in the conduction processes within plants. Sugars are conveyed throughout the plant in the phloem; water and other nutrients pass through the xylem.
The fascicular and interfascicular cambia thus join up to form a ring (in three dimensions, a tube) which separates the primary xylem and primary phloem, the cambium ring. The vascular cambium produces secondary xylem on the inside of the ring, and secondary phloem on the outside, pushing the primary xylem and phloem apart.
The phloem tissue arises from the cell facing outside and consists of sieve tubes and their companion cells. The function of phloem tissue is to distribute food from photosynthetic tissue to other tissues. The two tissues are separated by cambium, a tissue that divides to form xylem or phloem cells.
The earliest vascular plants had stems with a central core of vascular tissue. [3] [4] This consisted of a cylindrical strand of xylem, surrounded by a region of phloem. Around the vascular tissue there might have been an endodermis that regulated the flow of water into and out of the vascular system. Such an arrangement is termed a protostele.
Phloem was introduced by Carl Nägeli in 1858 after the discovery of sieve elements. Since then, multiple studies have been conducted on how sieve elements function in phloem in terms of working as a transport mechanism. [2] An example of analysis of phloem through sieve elements was conducted in the study of Arabidopsis leaves.