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The evolution of tetrapods began about 400 million years ago in the Devonian Period with the earliest tetrapods evolved from lobe-finned fishes. [1] Tetrapods (under the apomorphy-based definition used on this page) are categorized as animals in the biological superclass Tetrapoda, which includes all living and extinct amphibians, reptiles, birds, and mammals.
In effect, "tetrapod" is a name reserved solely for animals which lie among living tetrapods, so-called crown tetrapods. This is a node-based clade , a group with a common ancestry descended from a single "node" (the node being the nearest common ancestor of living species).
Early fossil tetrapods have been found in marine sediments, and because fossils of primitive tetrapods in general are found scattered all around the world, they must have spread by following the coastal lines — they could not have lived in freshwater only. Fossil Illuminates Evolution of Limbs from Fins Scientific American, 2 2 April 2004.
Tiktaalik, a lobe-finned fish with some anatomical features similar to early tetrapods. It has been suggested to be a transitional species between fish and tetrapods. [81] 365 Ma Acanthostega is one of the earliest vertebrates capable of walking. [82] 363 Ma By the start of the Carboniferous Period, the Earth begins to resemble its present state.
Acanthostega is a partially aquatic tetrapod with developed limbs that shares features common with the earlier tetrapods, Panderichthys and Eusthenopteron. [3] Like Panderichthys , the humerus of Acanthostega is flattened dorso-ventrally, the intermedium terminates level with the radius, and the endoskeleton can be divided into stylopodium ...
The fins of lobe-finned fish differ from those of all other fish in that each is borne on a fleshy, lobelike, scaly stalk extending from the body. The pectoral and pelvic fins are articulated in ways resembling the tetrapod limbs they were the precursors to. The fins evolved into the legs of the first tetrapod land vertebrates, amphibians.
Though stem-tetrapods originated in the preceding Devonian, it was in the earliest Carboniferous that the first crown tetrapods appeared, with full scaleless skin and five digits. During this time, amphibians (including many extinct groups unrelated to modern forms, referred to as "basal tetrapods") were the predominant tetrapods, and included ...
The tetrapods evolved from the lobe-finned fishes about 395 million years ago in the Devonian. [23] The specific aquatic ancestors of the tetrapods, and the process by which land colonization occurred, remain unclear, and are areas of active research and debate among palaeontologists at present.