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In trees, the phloem is the innermost layer of the bark, hence the name, derived from the Ancient Greek word φλοιός (phloiós), meaning "bark". [3] [4] The term was introduced by Carl Nägeli in 1858. [5] [6] Different types of phloem can be distinguished. The early phloem formed in the growth apices is called protophloem.
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The first xylem to develop is called 'protoxylem'. In appearance, protoxylem is usually distinguished by narrower vessels formed of smaller cells. Some of these cells have walls that contain thickenings in the form of rings or helices. Functionally, protoxylem can extend: the cells can grow in size and develop while a stem or root is elongating.
According to the hypothesis, the high concentration of organic substances, particularly sugar, inside the phloem at a source such as a leaf creates a diffusion gradient (osmotic gradient) that draws water into the cells from the adjacent xylem. This creates turgor pressure, also called hydrostatic pressure, in the phloem. The hypothesis states ...
Primarily, phloem carries dissolved food substances throughout the plant. This conduction system is composed of sieve-tube member and companion cells, that are without secondary walls. The parent cells of the vascular cambium produce both xylem and phloem. This usually also includes fibers, parenchyma and ray cells.
The phenolic unit can be found dimerized or further polymerized, creating a new class of polyphenol. For example, ellagic acid is a dimer of gallic acid and forms the class of ellagitannins, or a catechin and a gallocatechin can combine to form the red compound theaflavin , a process that also results in the large class of brown thearubigins in ...
Cross-section of a flax plant stem: 1. Pith 2. Protoxylem 3. Xylem I 4. Phloem I 5. Sclerenchyma 6. Cortex 7. Epidermis. In botany, a cortex is an outer layer of a stem or root in a vascular plant, lying below the epidermis but outside of the vascular bundles. [1]
An endodermis generally surrounds the stele. A centrarch (protoxylem in the center of a metaxylem cylinder) haplostele is prevalent in members of the rhyniophyte grade, such as Rhynia. [6] actinostele – a variation of the protostele in which the core is lobed or fluted. [7] This stele is found in many species of club moss (Lycopodium and ...