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A chordate (/ ˈ k ɔːr d eɪ t / KOR-dayt) is a deuterostomal bilaterian animal belonging to the phylum Chordata (/ k ɔːr ˈ d eɪ t ə / kor-DAY-tə).All chordates possess, at some point during their larval or adult stages, five distinctive physical characteristics (synapomorphies) that distinguish them from other taxa.
This article contains a list of all of the classes and orders that are located in the Phylum Chordata. The subphyla Tunicata and Vertebrata are in the unranked Olfactores clade, while the subphylum Cephalochordata is not. Animals in Olfactores are characterized as having a more advanced olfactory system than animals not in it.
Another feature present in both the Hemichordata and Chordata is pharyngotremy — the presence of spiracles or gill slits into the pharynx, which is also found in some primitive fossil echinoderms . [17] [18] A hollow nerve cord is found in all chordates, including tunicates (in the larval stage). Some hemichordates also have a tubular nerve cord.
The calcichordate hypothesis, formulated by British Museum paleontologist Richard Jefferies, holds that each separate lineage of chordate (Cephalochordates, Urochordates, Craniates) evolved from its own lineage of mitrate, and thus the echinoderms and the chordates are sister groups, with the hemichordates as an out-group. [1]
The Cambrian chordates are an extinct group of animals belonging to the phylum Chordata that lived during the Cambrian, between 538 and 485 million years ago. The first Cambrian chordate known is Pikaia gracilens , a lancelet -like animal from the Burgess Shale in British Columbia , Canada.
The notochord is the defining feature (synapomorphy) of chordates, and was present throughout life in many of the earliest chordates. Although the stomochord of hemichordates was once thought to be homologous or from a common lineal origin, it is now viewed as analogous, convergent, or from a different lineal origin. [22]
The Chordata ancestor gave rise to the lancelets (Amphioxii) and Olfactores. Ancestral chordates evolved a post-anal tail, notochord, and endostyle (precursor of thyroid). The pharyngeal slits (or gills) are now supported by connective tissue and used for filter feeding and possibly breathing.
Pikaia gracilens is an extinct, primitive chordate marine animal known from the Middle Cambrian Burgess Shale of British Columbia.Described in 1911 by Charles Doolittle Walcott as an annelid, and in 1979 by Harry B. Whittington and Simon Conway Morris as a chordate, it became "the most famous early chordate fossil", [1] or "famously known as the earliest described Cambrian chordate". [2]