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The precursor to the male angiosperm gametophyte is a diploid microspore mother cell located inside the anther. Once the microspore undergoes meiosis, 4 haploid cells are formed, each of which is a singled celled male gametophyte. The male gametophyte will develop via one or two rounds of mitosis inside the anther.
It is the biological process of gametogenesis during which cells that are haploid or diploid divide to create other cells. It can take place either through mitotic or meiotic division of diploid gametocytes into different cells depending on an organism's biological life cycle. For instance, gametophytes in plants undergo mitosis to produce gametes.
These microspore mother cells, also called microsporocytes, then undergo meiosis and become four microspore haploid cells. These new microspore cells then undergo mitosis and form a tube cell and a generative cell. The generative cell then undergoes mitosis one more time to form two male gametes, also called sperm.
The different stages of mitosis altogether define the mitotic phase (M phase) of a cell cycle—the division of the mother cell into two daughter cells genetically identical to each other. [3] The process of mitosis is divided into stages corresponding to the completion of one set of activities and the start of the next.
The pollen grains, which are the male gametophytes, are reduced to only a few cells (just three cells in many cases). Here the notion of two generations is less obvious; as Bateman & Dimichele say "sporophyte and gametophyte effectively function as a single organism". [8] The alternative term 'alternation of phases' may then be more appropriate ...
Four chambers (pollen sacs) lined with nutritive tapetal cells are visible by the time the microspores are produced. After meiosis, the haploid microspores undergo several changes: The microspore divides by mitosis producing two cells. The first of the cells (the generative cell) is small and is formed inside the second larger cell (the tube cell).
The eukaryotic cell cycle consists of four distinct phases: G 1 phase, S phase (synthesis), G 2 phase (collectively known as interphase) and M phase (mitosis and cytokinesis). M phase is itself composed of two tightly coupled processes: mitosis, in which the cell's nucleus divides, and cytokinesis, in which the cell's cytoplasm and cell membrane divides forming two daughter cells.
These spores then germinate and divide by mitosis to form a haploid multicellular phase, the gametophyte, which produces gametes directly by mitosis. This type of life cycle, involving alternation between two multicellular phases, the sexual haploid gametophyte and asexual diploid sporophyte, is known as alternation of generations.