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Chelicerata split from Mandibulata by the mid-Cambrian, as evidenced by stem-group chelicerates like Habeliida and Mollisonia present by this time. [2] The surviving marine species include the four species of xiphosurans (horseshoe crabs), and possibly the 1,300 species of pycnogonids (sea spiders), if the latter are indeed chelicerates.
However, a 2022 analysis recovered the monophyly of Merostomata, as opposed to a monophyletic Arachnida, with Xiphosura as the only modern representative in a derived position, indicating the convergence of several characteristics that supposedly united the arachnids.
The microscopic mite Lorryia formosa (). The mites are not a defined taxon, but is used for two distinct groups of arachnids, the Acariformes and the Parasitiformes.The phylogeny of the Acari has been relatively little studied, but molecular information from ribosomal DNA is being extensively used to understand relationships between groups.
The jumping spider Phidippus audax.The basal parts of the chelicerae are the two iridescent green mouthparts. The chelicerae (/ k ə ˈ l ɪ s ər iː /) are the mouthparts of the subphylum Chelicerata, an arthropod group that includes arachnids, horseshoe crabs, and sea spiders.
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[not seen, in Russian with English summary, English translation in Russian journal of zoology, 2: 409–413. (1998)] Lourenço, W.R.; Pham, D.-S. 2010: A remarkable new cave scorpion of the family Pseudochactidae Gromov (Chelicerata, Scorpiones) from Vietnam.
The subphylum Mandibulata constitutes one of the major subdivisions of the phylum Arthropoda, alongside Chelicerata. Mandibulates include the crustaceans , myriapods ( centipedes and millipedes ), and all true insects .
While adults live freely and are often found wandering about, searching for small animals and insect eggs for food, the larvae try to find a host to attach themselves to, often an insect like a grasshopper or fly, but also arachnids like harvestmen or spiders.