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The recently elucidated archaeal flagellum, or archaellum, is analogous—but not homologous—to the bacterial one. In addition to no sequence similarity being detected between the genes of the two systems, the archaeal flagellum appears to grow at the base rather than the tip, and is about 15 nanometers (nm) in diameter rather than 20.
[45] [46] However, unlike the bacterial flagellum archaellum has not shown to play a role in archaeal biofilm formation. [47] In archaeal biofilms, the only proposed function is thus far during the dispersal phase of biofilm when archaeal cells escape the community using their archaellum to further initiate the next round of biofilm formation.
The bacterial flagellum shares a common ancestor with the type III secretion system, [125] [126] while archaeal flagella appear to have evolved from bacterial type IV pili. [127] In contrast with the bacterial flagellum, which is hollow and assembled by subunits moving up the central pore to the tip of the flagella, archaeal flagella are ...
While bacterial cells often have many flagellar filaments, each of which rotates independently, the archaeal flagellum is composed of a bundle of many filaments that rotates as a single assembly. Bacterial flagella grow by the addition of flagellin subunits at the tip; archaeal flagella grow by the addition of subunits to the base.
[117] [118] [119] They provide two of several kinds of bacterial motility. [120] [121] Archaeal flagella are called archaella, and function in much the same way as bacterial flagella. Structurally the archaellum is superficially similar to a bacterial flagellum, but it differs in many details and is considered non-homologous. [122] [116]
Bacterial flagella are helical filaments, each with a rotary motor at its base which can turn clockwise or counterclockwise. [16] [17] [18] They provide two of several kinds of bacterial motility. [19] [20] Archaeal flagella are called archaella, and function in much the same way as bacterial flagella
The bacterial flagellum stretches from cytoplasm through the cell membrane(s) and extrudes through the cell wall. They are long and thick thread-like appendages, protein in nature. A different type of flagellum is found in archaea and a different type is found in eukaryotes.
Mitochondria and plastids contain their own ribosomes; these are more similar to those of bacteria (70S) than those of eukaryotes. [74] Proteins created by mitochondria and chloroplasts use N-formylmethionine as the initiating amino acid, as do proteins created by bacteria but not proteins created by eukaryotic nuclear genes or archaea. [75] [76]