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Schematic representation of a Euglena cell with red eyespot (9) Schematic representation of a Chlamydomonas cell with chloroplast eyespot (4). The eyespot apparatus (or stigma) is a photoreceptive organelle found in the flagellate or (motile) cells of green algae and other unicellular photosynthetic organisms such as euglenids.
Euglena is a genus of single cell flagellate eukaryotes. It is the best known and most widely studied member of the class Euglenoidea, a diverse group containing some 54 genera and at least 200 species. [1] [2] Species of Euglena are found in fresh water and salt water.
Euglena gracilis is a freshwater species of single-celled alga in the genus Euglena. It has secondary chloroplasts , and is a mixotroph able to feed by photosynthesis or phagocytosis . It has a highly flexible cell surface, allowing it to change shape from a thin cell up to 100 μm long to a sphere of approximately 20 μm.
The pigmented stigma is not to be confused with the photoreceptor. The stigma only provides directional shading for the adjacent membrane-inserted photoreceptors (the term "eyespot" is therefore misleading). Stigmata can also reflect and focus light like a concave mirror, thereby enhancing sensitivity. [57]
Euglenales consists mostly of freshwater organisms, in contrast to its sister Eutreptiales which is generally marine. Cells have two flagella, but only one is emergent; the other is very short and does not emerge from the cell, so cells appear to have only one flagellum. [3]
Euglenoids are distinguished mainly by the presence of a type of cell covering called a pellicle. Within its taxon, the pellicle is one of the euglenoids' most diverse morphological features. [7] The pellicle is composed of proteinaceous strips underneath the cell membrane, supported by dorsal and ventral microtubules. This varies from rigid to ...
Peranema cells are gliding flagellates found in freshwater lakes, ponds and ditches, and are often abundant at the bottom of stagnant pools rich in decaying organic material. [2] Although they belong to the class Euglenoidea , and are morphologically similar to the green Euglena , Peranema have no chloroplasts , and do not conduct autotrophy .
As with other euglenids, cells in the Euglenaceae are surrounded by a series of proteinaceous strips called the pellicle; the pellicle can stretch in most genera, allowing the cell to contract, creating a type of movement called metaboly.