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In 2011 the OEC of PSII was resolved to a level of 1.9Å revealing five oxygen atoms serving as oxo bridges linking the five metal atoms and four water molecules bound to the Mn 4 CaO 5 cluster; more than 1,300 water molecules were found in each photosystem II monomer, some forming extensive hydrogen-bonding networks that may serve as channels ...
The cyclic light-dependent reactions occur only when the sole photosystem being used is photosystem I. Photosystem I excites electrons which then cycle from the transport protein, ferredoxin (Fd), to the cytochrome complex, b 6 f, to another transport protein, plastocyanin (Pc), and back to photosystem I. A proton gradient is created across the ...
The antenna complex contains hundreds of chlorophyll molecules which funnel the excitation energy to the center of the photosystem. At the reaction center, the energy will be trapped and transferred to produce a high energy molecule. [2] The main function of PSII is to efficiently split water into oxygen molecules and protons.
An electron transport chain (ETC [1]) is a series of protein complexes and other molecules which transfer electrons from electron donors to electron acceptors via redox reactions (both reduction and oxidation occurring simultaneously) and couples this electron transfer with the transfer of protons (H + ions) across a membrane.
The antenna complex is composed of molecules of chlorophyll and carotenoids mounted on two proteins. [11] These pigment molecules transmit the resonance energy from photons when they become photoexcited. Antenna molecules can absorb all wavelengths of light within the visible spectrum. [12]
The evolution of oxygen during the light-dependent steps in photosynthesis (Hill reaction) was proposed and proven by British biochemist Robin Hill. He demonstrated that isolated chloroplasts would make oxygen (O 2) but not fix carbon dioxide (CO 2). This is evidence that the light and dark reactions occur at different sites within the cell. [1 ...
Manganese also forms strong bonds with oxygen-containing molecules such as water. The process of oxidizing two molecules of water to form an oxygen molecule requires four electrons. The water molecules that are oxidized in the manganese center are the source of the electrons that reduce the two molecules of Q to QH 2.
The last steps of this process occur in mitochondria. The reduced molecules NADH and FADH 2 are generated by the Krebs cycle, glycolysis, and pyruvate processing. These molecules pass electrons to an electron transport chain, which releases the energy of oxygen to create a proton gradient across the inner mitochondrial membrane.