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Palisade cells are located beneath the upper epidermis and cuticle but above the spongy mesophyll cells. Palisade cells contain a high concentration of chloroplasts , particularly in the upper portion of the cell, making them the primary site of photosynthesis in the leaves of plants that contain them.
In most leaves, the primary photosynthetic tissue is the palisade mesophyll and is located on the upper side of the blade or lamina of the leaf, [1] but in some species, including the mature foliage of Eucalyptus, [5] palisade mesophyll is present on both sides and the leaves are said to be isobilateral.
In leaves, they form two layers of mesophyll cells immediately beneath the epidermis of the leaf, that are responsible for photosynthesis and the exchange of gases. [2] These layers are called the palisade parenchyma and spongy mesophyll. Palisade parenchyma cells can be either cuboidal or elongated.
Loosely-arranged mesophyll cells lie between the bundle sheath and the leaf surface. The Calvin cycle is confined to the chloroplasts of these bundle sheath cells in C 4 plants. C 2 plants also use a variation of this structure. [1]
Spongy tissue is a type of tissue found both in plants and animals.. In plants, it is part of the mesophyll, where it forms a layer next to the palisade cells in the leaf.The spongy mesophyll's function is to allow for the interchange of gases (CO 2) that are needed for photosynthesis.
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Palisade mesophyll cells can contain 30–70 chloroplasts per cell, while stomatal guard cells contain only around 8–15 per cell, as well as much less chlorophyll. Chloroplasts can also be found in the bundle sheath cells of a leaf, especially in C 4 plants , which carry out the Calvin cycle in their bundle sheath cells.