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Chelicerata split from Mandibulata by the mid-Cambrian, as evidenced by stem-group chelicerates like Habeliida and Mollisonia present by this time. [2] The surviving marine species include the four species of xiphosurans (horseshoe crabs), and possibly the 1,300 species of pycnogonids (sea spiders), if the latter are indeed chelicerates. On the ...
The chelicerae (/ k ə ˈ l ɪ s ər iː /) are the mouthparts of the subphylum Chelicerata, an arthropod group that includes arachnids, horseshoe crabs, and sea spiders. Commonly referred to as "jaws", chelicerae may be shaped as either articulated fangs, or as a type of pincers.
Paired maxillae cut food and manipulate it during mastication. Maxillae can have hairs and "teeth" along their inner margins. At the outer margin, the galea is a cupped or scoop-like structure, which sits over the outer edge of the labium. They also have palps, which are used to sense the characteristics of potential foods.
The Crustacea were generally considered the closest relatives of the Uniramia, and sometimes these were united as Mandibulata.However, the competing hypothesis — that Crustacea and Hexapoda form a monophyletic group, the Pancrustacea, to which the Myriapoda are the closest relatives — has support from molecular and fossil evidence.
Harpactea sadistica is the first spider species – and the first member of the entire subphylum Chelicerata – found to use traumatic insemination. [2] The males have specialized genital structures at the pedipalps that are adapted to grip the female and inject the sperm, using a structure resembling a hypodermic needle.
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The subphylum Mandibulata constitutes one of the major subdivisions of the phylum Arthropoda, alongside Chelicerata. Mandibulates include the crustaceans, myriapods (centipedes and millipedes), and all true insects.
Sanctacaris is a Middle Cambrian arthropod from the Burgess Shale of British Columbia.It was most famously regarded as a stem-group chelicerate, a group which includes horseshoe crabs, spiders and scorpions, although subsequent phylogenetic studies have not always supported this conclusion. [1]