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The nucleolus (/ nj uː ˈ k l iː ə l ə s, ˌ nj uː k l i ˈ oʊ l ə s /; pl.: nucleoli /-l aɪ /) is the largest structure in the nucleus of eukaryotic cells. [1] It is best known as the site of ribosome biogenesis .
The cell nucleus (from Latin nucleus or nuculeus ' kernel, seed '; pl.: nuclei) is a membrane-bound organelle found in eukaryotic cells.Eukaryotic cells usually have a single nucleus, but a few cell types, such as mammalian red blood cells, have no nuclei, and a few others including osteoclasts have many.
The location of NORs and the nucleolar cycle in human cells. Nucleolus organizer regions (NORs) are chromosomal regions crucial for the formation of the nucleolus.In humans, the NORs are located on the short arms of the acrocentric chromosomes 13, 14, 15, 21 and 22, the genes RNR1, RNR2, RNR3, RNR4, and RNR5 respectively. [1]
The organization of chromosomes into distinct regions within the nucleus was first proposed in 1885 by Carl Rabl.Later in 1909, with the help of the microscopy technology at the time, Theodor Boveri coined the termed chromosome territories after observing that chromosomes occupy individually distinct nuclear regions. [6]
Some nucleoporins which typically make up the nuclear pore, can be mobile and participate in the regulation of gene expression in the nucleoplasm. [8] [9] The nuclear pore is where molecules travel from inside the nucleoplasm to the cytoplasm and vice versa. [9] The nucleoplasm is also a route for many molecules to travel through. [9]
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Binucleated cells can be observed using microscopy. Cells must first be fixed to arrest them wherever they are in the cell cycle and to keep their structures from degrading. Their nuclei and tubulin must next be made visible so that binucleation can be identified. DAPI is a dye that binds to DNA and fluoresces blue. For this reason, it is ...