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However, their ability to extract energy from cellulose digestion is less efficient than in ruminants. [2] Herbivores digest cellulose by microbial fermentation. Monogastric herbivores which can digest cellulose nearly as well as ruminants are called hindgut fermenters, while ruminants are called foregut fermenters. [3]
Ribbon representation of the Streptomyces lividans β-1,4-endoglucanase catalytic domain - an example from the family 12 glycoside hydrolases [1]. Cellulase (EC 3.2.1.4; systematic name 4-β-D-glucan 4-glucanohydrolase) is any of several enzymes produced chiefly by fungi, bacteria, and protozoans that catalyze cellulolysis, the decomposition of cellulose and of some related polysaccharides:
Some animals, particularly ruminants and termites, can digest cellulose with the help of symbiotic micro-organisms that live in their guts, such as Trichonympha. In human nutrition , cellulose is a non-digestible constituent of insoluble dietary fiber , acting as a hydrophilic bulking agent for feces and potentially aiding in defecation .
The most well-known of these indigestible compounds is cellulose; the basic chemical polymer in the makeup of plant cell walls. Most animals, however, do not produce cellulase; the enzyme needed to digest cellulose. However, some animals and species have developed symbiotic relationships with cellulase-producing bacteria (see termites and ...
Fermentation is crucial to digestion because it breaks down complex carbohydrates, such as cellulose, and enables the animal to use them. Microbes function best in a warm, moist, anaerobic environment with a temperature range of 37.7 to 42.2 °C (99.9 to 108.0 °F) and a pH between 6.0 and 6.4.
Cellulose is a polymer made with repeated glucose units bonded together by beta-linkages. Humans and many animals lack an enzyme to break the beta-linkages, so they do not digest cellulose. Certain animals, such as termites can digest cellulose, because bacteria possessing the enzyme are present in their gut. Cellulose is insoluble in water.
In the rumen, anaerobic organisms, including methanogens, digest cellulose into forms nutritious to the animal. Without these microorganisms, animals such as cattle would not be able to consume grasses. The useful products of methanogenesis are absorbed by the gut, but methane is released from the animal mainly by belching (eructation).
Mature leaves contain a high proportion of hard-to-digest cellulose, less energy than other types of foods, and often toxic compounds. [1] For this reason, folivorous animals tend to have long digestive tracts and slow metabolisms. Many enlist the help of symbiotic bacteria to release the nutrients in their diet.