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However, the punctuational equilibrium model may still be inferred from both the observation of stasis and examples of rapid and episodic speciation events documented in the fossil record. [70] Dawkins also emphasizes that punctuated equilibrium has been "oversold by some journalists", [71] but partly due to Eldredge and Gould's "later writings ...
The Bak–Sneppen model is a simple model of co-evolution between interacting species. It was developed to show how self-organized criticality may explain key features of the fossil record, such as the distribution of sizes of extinction events and the phenomenon of punctuated equilibrium. It is named after Per Bak and Kim Sneppen.
Dawkins could, should, and probably would accept it; in The Ancestor's Tale, he has an inclusive view of speciation mechanisms." (p. 100) Thus, while "Gould somewhat overstates the adherence of orthodoxy to strict extrapolationism", punctuated equilibrium is more important than some of the more "ungenerous treatment" that has been meted out.
Eldredge and Stephen Jay Gould proposed punctuated equilibria in 1972. Punctuated equilibrium is a refinement to evolutionary theory. It describes patterns of descent taking place in "fits and starts" separated by long periods of stability. Eldredge went on to develop a hierarchical vision of evolutionary and ecological systems. Around this ...
In the 1980s, the American palaeontologists Stephen Jay Gould and Niles Eldredge argued for an extended synthesis based on their idea of punctuated equilibrium, the role of species selection shaping large scale evolutionary patterns and natural selection working on multiple levels extending from genes to species. [30] [31] [32] [33]
Gould's most significant contribution to evolutionary biology was the theory of punctuated equilibrium [2] developed with Niles Eldredge in 1972. [3] The theory proposes that most evolution is characterized by long periods of evolutionary stability, infrequently punctuated by swift periods of branching speciation.
While the traditional model of paleontology, the phylogenetic model, posits that features evolved slowly without any direct association with speciation, the relatively newer and more controversial idea of punctuated equilibrium claims that major evolutionary changes don't happen over a gradual period but in localized, rare, rapid events of branching speciation.
The theory of punctuated equilibrium developed by Stephen Jay Gould and Niles Eldredge and first presented in 1972 [61] is often mistakenly drawn into the discussion of transitional fossils. [62] This theory, however, pertains only to well-documented transitions within taxa or between closely related taxa over a geologically short period of time.