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However, the punctuational equilibrium model may still be inferred from both the observation of stasis and examples of rapid and episodic speciation events documented in the fossil record. [70] Dawkins also emphasizes that punctuated equilibrium has been "oversold by some journalists", [71] but partly due to Eldredge and Gould's "later writings ...
Dawkins could, should, and probably would accept it; in The Ancestor's Tale, he has an inclusive view of speciation mechanisms." (p. 100) Thus, while "Gould somewhat overstates the adherence of orthodoxy to strict extrapolationism", punctuated equilibrium is more important than some of the more "ungenerous treatment" that has been meted out.
The Bak–Sneppen model is a simple model of co-evolution between interacting species. It was developed to show how self-organized criticality may explain key features of the fossil record, such as the distribution of sizes of extinction events and the phenomenon of punctuated equilibrium. It is named after Per Bak and Kim Sneppen.
Gould's most significant contribution to evolutionary biology was the theory of punctuated equilibrium [2] developed with Niles Eldredge in 1972. [3] The theory proposes that most evolution is characterized by long periods of evolutionary stability, infrequently punctuated by swift periods of branching speciation.
There are four geographic modes of speciation in nature, based on the extent to which speciating populations are isolated from one another: allopatric, peripatric, parapatric, and sympatric. Whether genetic drift is a minor or major contributor to speciation is the subject of much ongoing discussion.
The theory of punctuated equilibrium developed by Stephen Jay Gould and Niles Eldredge and first presented in 1972 [61] is often mistakenly drawn into the discussion of transitional fossils. [62] This theory, however, pertains only to well-documented transitions within taxa or between closely related taxa over a geologically short period of time.
Gould's initial work on punctuated equilibrium was coauthored with Niles Eldredge. [20] Gould's model of punctuated equilibrium drew attention to episodic bursts of evolutionary change followed by periods of morphological stability. He challenged the conventional model of gradual, continuous change - called phyletic gradualism. [21]
Two useful introductions to the fundamental theory underlying the unit of selection issue and debate, which also present examples of multi-level selection from the entire range of the biological hierarchy (typically with entities at level N-1 competing for increased representation, i.e., higher frequency, at the immediately higher level N, e.g., organisms in populations or cell lineages in ...